UniProtKB - Q9Y5J1 (UTP18_HUMAN)

BLAST

>sp|Q9Y5J1|UTP18_HUMAN U3 small nucleolar RNA-associated protein 18 homolog OS=Homo sapiens OX=9606 GN=UTP18 PE=1 SV=3
MPPERRRRMKLDRRTGAKPKRKPGMRPDWKAGAGPGGPPQKPAPSSQRKPPARPSAAAAA
IAVAAAEEERRLRQRNRLRLEEDKPAVERCLEELVFGDVENDEDALLRRLRGPRVQEHED
SGDSEVENEAKGNFPPQKKPVWVDEEDEDEEMVDMMNNRFRKDMMKNASESKLSKDNLKK
RLKEEFQHAMGGVPAWAETTKRKTSSDDESEEDEDDLLQRTGNFISTSTSLPRGILKMKN
CQHANAERPTVARISSVQFHPGAQIVMVAGLDNAVSLFQVDGKTNPKIQSIYLERFPIFK
ACFSANGEEVLATSTHSKVLYVYDMLAGKLIPVHQVRGLKEKIVRSFEVSPDGSFLLING
IAGYLHLLAMKTKELIGSMKINGRVAASTFSSDSKKVYASSGDGEVYVWDVNSRKCLNRF
VDEGSLYGLSIATSRNGQYVACGSNCGVVNIYNQDSCLQETNPKPIKAIMNLVTGVTSLT
FNPTTEILAIASEKMKEAVRLVHLPSCTVFSNFPVIKNKNISHVHTMDFSPRSGYFALGN
EKGKALMYRLHHYSDF

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History
Entry version 174 (07 Nov 2018)
Sequence version 3 (30 Aug 2005)
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Protein

U3 small nucleolar RNA-associated protein 18 homolog

Gene

UTP18

Organism

Homo sapiens (Human)

Status

Reviewed-Annotation score: -Experimental evidence at protein levelⁱ

Functionⁱ

Involved in nucleolar processing of pre-18S ribosomal RNA.By similarity

GO - Molecular functionⁱ

RNA binding
Source: UniProtKB
Inferred from high throughput direct assayⁱ
- 22658674
- 22681889

View the complete GO annotation on QuickGO ...

GO - Biological processⁱ

rRNA processing Source: Reactome

View the complete GO annotation on QuickGO ...

Keywordsⁱ

Biological process	rRNA processing

Biological process

rRNA processing

Enzyme and pathway databases

Reactomeⁱ	R-HSA-6790901 rRNA modification in the nucleus and cytosol R-HSA-6791226 Major pathway of rRNA processing in the nucleolus and cytosol

Reactomeⁱ

R-HSA-6790901 rRNA modification in the nucleus and cytosol
R-HSA-6791226 Major pathway of rRNA processing in the nucleolus and cytosol

Names & Taxonomyⁱ

Protein namesⁱ	Recommended name: U3 small nucleolar RNA-associated protein 18 homolog Alternative name(s): WD repeat-containing protein 50
Gene namesⁱ	Name:UTP18 Synonyms:WDR50 ORF Names:CDABP0061, CGI-48
Organismⁱ	Homo sapiens (Human)
Taxonomic identifierⁱ	9606 [NCBI]
Taxonomic lineageⁱ	cellular organisms › Eukaryota › Opisthokonta › Metazoa › Eumetazoa › Bilateria › Deuterostomia › Chordata › Craniata › Vertebrata › Gnathostomata › Teleostomi › Euteleostomi › Sarcopterygii › Dipnotetrapodomorpha › Tetrapoda › Amniota › Mammalia › Theria › Eutheria › Boreoeutheria › Euarchontoglires › Primates › Haplorrhini › Simiiformes › Catarrhini › Hominoidea › Hominidae › Homininae › Homo
Proteomesⁱ	UP000005640 Componentⁱ: Chromosome 17

Organism-specific databases

EuPathDBⁱ	HostDB:ENSG00000011260.13
Human Gene Nomenclature Database More...HGNCⁱ	HGNC:24274 UTP18
MIMⁱ	612816 gene
neXtProtⁱ	NX_Q9Y5J1

Keywords - Cellular componentⁱ

Nucleus

Pathology & Biotechⁱ

Organism-specific databases

DisGeNET More...DisGeNETⁱ	51096
Open Targets More...OpenTargetsⁱ	ENSG00000011260
PharmGKBⁱ	PA134984104

Polymorphism and mutation databases

BioMutaⁱ	UTP18
DMDMⁱ	73920973

PTM / Processingⁱ

Molecule processing

Feature key	Position(s)	DescriptionActions	Graphical view	Length
ChainⁱPRO_0000051405	1 – 556	U3 small nucleolar RNA-associated protein 18 homologAdd BLAST		556

Amino acid modifications

Feature key	Position(s)	DescriptionActions	Length
Cross-linkⁱ	84	Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2)Combined sources
Modified residueⁱ	121	PhosphoserineCombined sources	1
Modified residueⁱ	124	PhosphoserineCombined sources	1
Cross-linkⁱ	183	Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2)Combined sources
Cross-linkⁱ	201	Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2)Combined sources
Modified residueⁱ	204	PhosphothreonineCombined sources	1
Modified residueⁱ	205	PhosphoserineCombined sources	1
Modified residueⁱ	206	PhosphoserineCombined sources	1
Modified residueⁱ	210	PhosphoserineCombined sources	1
Modified residueⁱ	221	PhosphothreonineCombined sources	1
Cross-linkⁱ	517	Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2)Combined sources

Keywords - PTMⁱ

Isopeptide bond, Phosphoprotein, Ubl conjugation

Proteomic databases

EPDⁱ	Q9Y5J1
MaxQB - The MaxQuant DataBase More...MaxQBⁱ	Q9Y5J1
PaxDbⁱ	Q9Y5J1
PeptideAtlas More...PeptideAtlasⁱ	Q9Y5J1
PRIDEⁱ	Q9Y5J1
ProteomicsDBⁱ	86417

2D gel databases

SWISS-2DPAGEⁱ	Q9Y5J1

SWISS-2DPAGEⁱ

Q9Y5J1

PTM databases

iPTMnetⁱ	Q9Y5J1
PhosphoSitePlusⁱ	Q9Y5J1

Miscellaneous databases

PMAP-CutDBⁱ	Q9Y5J1

PMAP-CutDBⁱ

Q9Y5J1

Expressionⁱ

Gene expression databases

Bgeeⁱ	ENSG00000011260 Expressed in 231 organ(s), highest expression level in testis
CleanExⁱ	HS_UTP18
ExpressionAtlasⁱ	Q9Y5J1 baseline and differential
Genevisibleⁱ	Q9Y5J1 HS

Organism-specific databases

Human Protein Atlas More...HPAⁱ	HPA052378

HPAⁱ

HPA052378

Interactionⁱ

Protein-protein interaction databases

BioGridⁱ	119285, 30 interactors
IntActⁱ	Q9Y5J1, 17 interactors
Molecular INTeraction database More...MINTⁱ	Q9Y5J1
STRINGⁱ	9606.ENSP00000225298

Structureⁱ

3D structure databases

ProteinModelPortalⁱ	Q9Y5J1
SMRⁱ	Q9Y5J1
ModBaseⁱ	Search...
MobiDBⁱ	Search...

Family & Domainsⁱ

Domains and Repeats

Feature key	Position(s)	DescriptionActions	Length
Repeatⁱ	249 – 288	WD 1Add BLAST	40
Repeatⁱ	293 – 333	WD 2Add BLAST	41
Repeatⁱ	339 – 380	WD 3Add BLAST	42
Repeatⁱ	381 – 419	WD 4Add BLAST	39
Repeatⁱ	421 – 462	WD 5Add BLAST	42
Repeatⁱ	471 – 512	WD 6Add BLAST	42

Sequence similaritiesⁱ

Belongs to the WD repeat UTP18 family.Curated

Keywords - Domainⁱ

Repeat, WD repeat

Phylogenomic databases

eggNOGⁱ	KOG2055 Eukaryota ENOG410YBWP LUCA
Ensembl GeneTree More...GeneTreeⁱ	ENSGT00440000033919
HOGENOMⁱ	HOG000242429
HOVERGENⁱ	HBG082494
InParanoidⁱ	Q9Y5J1
KEGG Orthology (KO) More...KOⁱ	K14553
OMAⁱ	QFHPTST
Database of Orthologous Groups More...OrthoDBⁱ	EOG091G0BJP
PhylomeDBⁱ	Q9Y5J1
TreeFamⁱ	TF313426

Family and domain databases

Gene3Dⁱ	2.130.10.10, 1 hit
InterProⁱ	View protein in InterPro IPR015943 WD40/YVTN_repeat-like_dom_sf IPR001680 WD40_repeat IPR019775 WD40_repeat_CS IPR017986 WD40_repeat_dom IPR036322 WD40_repeat_dom_sf
SMARTⁱ	View protein in SMART SM00320 WD40, 7 hits
SUPFAMⁱ	SSF50978 SSF50978, 1 hit
PROSITEⁱ	View protein in PROSITE PS00678 WD_REPEATS_1, 1 hit PS50082 WD_REPEATS_2, 1 hit PS50294 WD_REPEATS_REGION, 1 hit

Sequence (1+)ⁱ

Sequence statusⁱ: Complete.

This entry has 1 described isoform and 5 potential isoforms that are computationally mapped.Show all Align All

Q9Y5J1-1 [UniParc]FASTA Add to basket

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        10         20         30         40         50
MPPERRRRMK LDRRTGAKPK RKPGMRPDWK AGAGPGGPPQ KPAPSSQRKP 
        60         70         80         90        100
PARPSAAAAA IAVAAAEEER RLRQRNRLRL EEDKPAVERC LEELVFGDVE 
       110        120        130        140        150
NDEDALLRRL RGPRVQEHED SGDSEVENEA KGNFPPQKKP VWVDEEDEDE 
       160        170        180        190        200
EMVDMMNNRF RKDMMKNASE SKLSKDNLKK RLKEEFQHAM GGVPAWAETT 
       210        220        230        240        250
KRKTSSDDES EEDEDDLLQR TGNFISTSTS LPRGILKMKN CQHANAERPT 
       260        270        280        290        300
VARISSVQFH PGAQIVMVAG LDNAVSLFQV DGKTNPKIQS IYLERFPIFK 
       310        320        330        340        350
ACFSANGEEV LATSTHSKVL YVYDMLAGKL IPVHQVRGLK EKIVRSFEVS 
       360        370        380        390        400
PDGSFLLING IAGYLHLLAM KTKELIGSMK INGRVAASTF SSDSKKVYAS 
       410        420        430        440        450
SGDGEVYVWD VNSRKCLNRF VDEGSLYGLS IATSRNGQYV ACGSNCGVVN 
       460        470        480        490        500
IYNQDSCLQE TNPKPIKAIM NLVTGVTSLT FNPTTEILAI ASEKMKEAVR 
       510        520        530        540        550
LVHLPSCTVF SNFPVIKNKN ISHVHTMDFS PRSGYFALGN EKGKALMYRL 

HHYSDF

Length:556

Mass (Da):62,004

Last modified:August 30, 2005 - v3

Checksum:ⁱ800139F974F4EDF6

Computationally mapped potential isoform sequencesⁱ

There are 5 potential isoforms mapped to this entry.BLAST Align Show all Add to basket

Entry	Entry name	Protein names	Gene names	Length	Annotation

J3QR85	J3QR85_HUMAN	U3 small nucleolar RNA-associated p... U3 small nucleolar RNA-associated protein 18 homolog	UTP18	143	Annotation score:
J3KSR7	J3KSR7_HUMAN	U3 small nucleolar RNA-associated p... U3 small nucleolar RNA-associated protein 18 homolog	UTP18	177	Annotation score:
J3QRG0	J3QRG0_HUMAN	U3 small nucleolar RNA-associated p... U3 small nucleolar RNA-associated protein 18 homolog	UTP18	123	Annotation score:
J3QLD6	J3QLD6_HUMAN	U3 small nucleolar RNA-associated p... U3 small nucleolar RNA-associated protein 18 homolog	UTP18	115	Annotation score:
F6VJE8	F6VJE8_HUMAN	U3 small nucleolar RNA-associated p... U3 small nucleolar RNA-associated protein 18 homolog	UTP18	43	Annotation score:

Sequence cautionⁱ

The sequence AAD34043 differs from that shown. Reason: Frameshift at position 19.Curated

The sequence AAG01999 differs from that shown. Reason: Erroneous initiation.Curated

Experimental Info

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Sequence conflictⁱ	543 – 556	GKALM…HYSDF → ARP in AAD34043 (PubMed:10810093).CuratedAdd BLAST		14

Sequence databases

Select the link destinations: EMBLⁱ GenBankⁱ DDBJⁱ Links Updated	AF151806 mRNA Translation: AAD34043.1 Frameshift. BC025276 mRNA Translation: AAH25276.2 AY007138 mRNA Translation: AAG01999.1 Different initiation.
The Consensus CDS (CCDS) project More...CCDSⁱ	CCDS42362.1
NCBI Reference Sequences More...RefSeqⁱ	NP_057085.2, NM_016001.2
UniGeneⁱ	Hs.709327

Genome annotation databases

Ensemblⁱ	ENST00000225298; ENSP00000225298; ENSG00000011260
GeneIDⁱ	51096
KEGGⁱ	hsa:51096
UCSC genome browser More...UCSCⁱ	uc002its.4 human

Similar proteinsⁱ

90% Identity
50% Identity

Protein	Similar proteins		Species	Score	Length	Source
Q9Y5J1	Uncharacterized protein		RHIRO		596	UniRef90_Q9Y5J1
Uncharacterized protein		RHIBE		585
UTP18 isoform 6 (Fragment)		PONAB		42
UTP18 isoform 6 (Fragment)		PANTR		42
U3 small nucleolar RNA-associated protein 18 homolog isoform X1		TARSY		580
+83

Protein	Similar proteins		Species	Score	Length	Source
Q9Y5J1	U3 small nucleolar RNA-associated protein 18 homolog		MOUSE		552	UniRef50_Q9Y5J1
Uncharacterized protein		RHIRO		596
Uncharacterized protein		RHIBE		585
UTP18 isoform 6 (Fragment)		PONAB		42
UTP18 isoform 6 (Fragment)		PANTR		42
+298

Cross-referencesⁱ

Sequence databases

Select the link destinations: EMBLⁱ GenBankⁱ DDBJⁱ Links Updated	AF151806 mRNA Translation: AAD34043.1 Frameshift. BC025276 mRNA Translation: AAH25276.2 AY007138 mRNA Translation: AAG01999.1 Different initiation.
CCDSⁱ	CCDS42362.1
RefSeqⁱ	NP_057085.2, NM_016001.2
UniGeneⁱ	Hs.709327

3D structure databases

ProteinModelPortalⁱ	Q9Y5J1
SMRⁱ	Q9Y5J1
ModBaseⁱ	Search...
MobiDBⁱ	Search...

Protein-protein interaction databases

BioGridⁱ	119285, 30 interactors
IntActⁱ	Q9Y5J1, 17 interactors
MINTⁱ	Q9Y5J1
STRINGⁱ	9606.ENSP00000225298

PTM databases

iPTMnetⁱ	Q9Y5J1
PhosphoSitePlusⁱ	Q9Y5J1

Polymorphism and mutation databases

BioMutaⁱ	UTP18
DMDMⁱ	73920973

2D gel databases

SWISS-2DPAGEⁱ	Q9Y5J1

SWISS-2DPAGEⁱ

Q9Y5J1

Proteomic databases

EPDⁱ	Q9Y5J1
MaxQBⁱ	Q9Y5J1
PaxDbⁱ	Q9Y5J1
PeptideAtlasⁱ	Q9Y5J1
PRIDEⁱ	Q9Y5J1
ProteomicsDBⁱ	86417

Protocols and materials databases

Structural Biology Knowledgebase	Search...

Structural Biology Knowledgebase

Search...

Genome annotation databases

Ensemblⁱ	ENST00000225298; ENSP00000225298; ENSG00000011260
GeneIDⁱ	51096
KEGGⁱ	hsa:51096
UCSCⁱ	uc002its.4 human

Organism-specific databases

CTDⁱ	51096
DisGeNETⁱ	51096
EuPathDBⁱ	HostDB:ENSG00000011260.13
GeneCardsⁱ	UTP18
HGNCⁱ	HGNC:24274 UTP18
HPAⁱ	HPA052378
MIMⁱ	612816 gene
neXtProtⁱ	NX_Q9Y5J1
OpenTargetsⁱ	ENSG00000011260
PharmGKBⁱ	PA134984104
GenAtlas: human gene database More...GenAtlasⁱ	Search...

Phylogenomic databases

eggNOGⁱ	KOG2055 Eukaryota ENOG410YBWP LUCA
GeneTreeⁱ	ENSGT00440000033919
HOGENOMⁱ	HOG000242429
HOVERGENⁱ	HBG082494
InParanoidⁱ	Q9Y5J1
KOⁱ	K14553
OMAⁱ	QFHPTST
OrthoDBⁱ	EOG091G0BJP
PhylomeDBⁱ	Q9Y5J1
TreeFamⁱ	TF313426

Enzyme and pathway databases

Reactomeⁱ	R-HSA-6790901 rRNA modification in the nucleus and cytosol R-HSA-6791226 Major pathway of rRNA processing in the nucleolus and cytosol

Reactomeⁱ

R-HSA-6790901 rRNA modification in the nucleus and cytosol
R-HSA-6791226 Major pathway of rRNA processing in the nucleolus and cytosol

Miscellaneous databases

ChiTaRSⁱ	UTP18 human
GeneWikiⁱ	UTP18
GenomeRNAiⁱ	51096
PMAP-CutDBⁱ	Q9Y5J1
Protein Ontology More...PROⁱ	PR:Q9Y5J1
SOURCEⁱ	Search...

Gene expression databases

Bgeeⁱ	ENSG00000011260 Expressed in 231 organ(s), highest expression level in testis
CleanExⁱ	HS_UTP18
ExpressionAtlasⁱ	Q9Y5J1 baseline and differential
Genevisibleⁱ	Q9Y5J1 HS

Family and domain databases

Gene3Dⁱ	2.130.10.10, 1 hit
InterProⁱ	View protein in InterPro IPR015943 WD40/YVTN_repeat-like_dom_sf IPR001680 WD40_repeat IPR019775 WD40_repeat_CS IPR017986 WD40_repeat_dom IPR036322 WD40_repeat_dom_sf
SMARTⁱ	View protein in SMART SM00320 WD40, 7 hits
SUPFAMⁱ	SSF50978 SSF50978, 1 hit
PROSITEⁱ	View protein in PROSITE PS00678 WD_REPEATS_1, 1 hit PS50082 WD_REPEATS_2, 1 hit PS50294 WD_REPEATS_REGION, 1 hit
ProtoNetⁱ	Search...

Entry informationⁱ

Entry nameⁱ	UTP18_HUMAN
Accessionⁱ	Q9Y5J1Primary (citable) accession number: Q9Y5J1 Secondary accession number(s): Q9H4N6
Entry historyⁱ	Integrated into UniProtKB/Swiss-Prot:	November 16, 2001
	Last sequence update:	August 30, 2005
	Last modified:	November 7, 2018
	This is version 174 of the entry and version 3 of the sequence. See complete history.
Entry statusⁱ	Reviewed (UniProtKB/Swiss-Prot)
Annotation program	Chordata Protein Annotation Program
Disclaimer	Any medical or genetic information present in this entry is provided for research, educational and informational purposes only. It is not in any way intended to be used as a substitute for professional medical advice, diagnosis, treatment or care.

Miscellaneousⁱ

Keywords - Technical termⁱ

Complete proteome, Reference proteome

Documents

SIMILARITY comments
Index of protein domains and families
MIM cross-references
Online Mendelian Inheritance in Man (MIM) cross-references in UniProtKB/Swiss-Prot
Human chromosome 17
Human chromosome 17: entries, gene names and cross-references to MIM

UniProtKB

UniParc

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UniRef

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Supporting data

UniProtKB - Q9Y5J1 (UTP18_HUMAN)

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U3 small nucleolar RNA-associated protein 18 homolog

UTP18

Select a section on the left to see content.

<p>This section provides any useful information about the protein, mostly biological knowledge.<p><a href='/help/function_section' target='_top'>More...</a></p>Functioni

<p>The <a href="http://www.geneontology.org/">Gene Ontology (GO)</a> project provides a set of hierarchical controlled vocabulary split into 3 categories:<p><a href='/help/gene_ontology' target='_top'>More...</a></p>GO - Molecular functioni

GO - Biological processi

Enzyme and pathway databases

<p>This section provides information about the protein and gene name(s) and synonym(s) and about the organism that is the source of the protein sequence.<p><a href='/help/names_and_taxonomy_section' target='_top'>More...</a></p>Names & Taxonomyi

Organism-specific databases

<p>This section provides information on the location and the topology of the mature protein in the cell.<p><a href='/help/subcellular_location_section' target='_top'>More...</a></p>Subcellular locationi

Nucleus

Nucleus

Other locations

Keywords - Cellular componenti

<p>This section provides information on the disease(s) and phenotype(s) associated with a protein.<p><a href='/help/pathology_and_biotech_section' target='_top'>More...</a></p>Pathology & Biotechi

Organism-specific databases

Polymorphism and mutation databases

<p>This section describes post-translational modifications (PTMs) and/or processing events.<p><a href='/help/ptm_processing_section' target='_top'>More...</a></p>PTM / Processingi

Molecule processing

Amino acid modifications

Keywords - PTMi

Proteomic databases

2D gel databases

PTM databases

Miscellaneous databases

<p>This section provides information on the expression of a gene at the mRNA or protein level in cells or in tissues of multicellular organisms.<p><a href='/help/expression_section' target='_top'>More...</a></p>Expressioni

Gene expression databases

Organism-specific databases

<p>This section provides information on the quaternary structure of a protein and on interaction(s) with other proteins or protein complexes.<p><a href='/help/interaction_section' target='_top'>More...</a></p>Interactioni

Protein-protein interaction databases

<p>This section provides information on the tertiary and secondary structure of a protein.<p><a href='/help/structure_section' target='_top'>More...</a></p>Structurei

3D structure databases

<p>This section provides information on sequence similarities with other proteins and the domain(s) present in a protein.<p><a href='/help/family_and_domains_section' target='_top'>More...</a></p>Family & Domainsi

Domains and Repeats

<p>This subsection of the ‘Family and domains’ section provides information about the sequence similarity with other proteins.<p><a href='/help/sequence_similarities' target='_top'>More...</a></p>Sequence similaritiesi

Keywords - Domaini

Phylogenomic databases

Family and domain databases

<p>This subsection of the ‘Sequence’ section reports difference(s) between the protein sequence shown in the UniProtKB entry and other available protein sequences derived from the same gene.<p><a href='/help/sequence_caution' target='_top'>More...</a></p>Sequence cautioni

Experimental Info

Sequence databases

Genome annotation databases

<p>This section is used to point to information related to entries and found in data collections other than UniProtKB.<p><a href='/help/cross_references_section' target='_top'>More...</a></p>Cross-referencesi

Sequence databases

3D structure databases

Protein-protein interaction databases

PTM databases

Polymorphism and mutation databases

2D gel databases

Proteomic databases

Protocols and materials databases

Genome annotation databases

Organism-specific databases

Phylogenomic databases

Enzyme and pathway databases

Miscellaneous databases

Gene expression databases

Family and domain databases

<p>This section provides general information on the entry.<p><a href='/help/entry_information_section' target='_top'>More...</a></p>Entry informationi

<p>This section contains any relevant information that doesn’t fit in any other defined sections<p><a href='/help/miscellaneous_section' target='_top'>More...</a></p>Miscellaneousi

Keywords - Technical termi

Documents

UniProt is an ELIXIR core data resource

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Functionⁱ

GO - Molecular functionⁱ

GO - Biological processⁱ

Names & Taxonomyⁱ

Subcellular locationⁱ

Keywords - Cellular componentⁱ

Pathology & Biotechⁱ

PTM / Processingⁱ

Keywords - PTMⁱ

Expressionⁱ

Interactionⁱ

Structureⁱ

Family & Domainsⁱ

Sequence similaritiesⁱ

Keywords - Domainⁱ

Sequence cautionⁱ

Cross-referencesⁱ

Entry informationⁱ

Miscellaneousⁱ

Keywords - Technical termⁱ