FRMD4A

Functionⁱ

Scaffolding protein that regulates epithelial cell polarity by connecting ARF6 activation with the PAR3 complex (By similarity). Plays a redundant role with FRMD4B in epithelial polarization (By similarity). May regulate MAPT secretion by activating ARF6-signaling (PubMed:27044754).By similarity1 Publication

Caution

It is uncertain whether Met-1 or Met-16 is the initiator.Curated

GO - Molecular functionⁱ

protein binding, bridging Source: Ensembl

View the complete GO annotation on QuickGO ...

GO - Biological processⁱ

establishment of epithelial cell polarity Source: Ensembl

View the complete GO annotation on QuickGO ...

Names & Taxonomyⁱ

Protein namesⁱ	Recommended name: FERM domain-containing protein 4A
Gene namesⁱ	Name:FRMD4AImported Synonyms:FRMD4, KIAA1294
Organismⁱ	Homo sapiens (Human)
Taxonomic identifierⁱ	9606 [NCBI]
Taxonomic lineageⁱ	cellular organisms › Eukaryota › Opisthokonta › Metazoa › Eumetazoa › Bilateria › Deuterostomia › Chordata › Craniata › Vertebrata › Gnathostomata › Teleostomi › Euteleostomi › Sarcopterygii › Dipnotetrapodomorpha › Tetrapoda › Amniota › Mammalia › Theria › Eutheria › Boreoeutheria › Euarchontoglires › Primates › Haplorrhini › Simiiformes › Catarrhini › Hominoidea › Hominidae › Homininae › Homo
Proteomesⁱ	UP000005640 Componentⁱ: Chromosome 10

Organism-specific databases

EuPathDBⁱ	HostDB:ENSG00000151474.19
Human Gene Nomenclature Database More...HGNCⁱ	HGNC:25491 FRMD4A
MIMⁱ	616305 gene
neXtProtⁱ	NX_Q9P2Q2

Keywords - Cellular componentⁱ

Cell junction, Cytoplasm, Cytoskeleton, Tight junction

Pathology & Biotechⁱ

Involvement in diseaseⁱ

Agenesis of the corpus callosum, with facial anomalies and cerebellar ataxia (CCAFCA)1 Publication

The disease is caused by mutations affecting the gene represented in this entry.

Disease descriptionAn autosomal recessive intellectual disability syndrome characterized by congenital microcephaly, low anterior hairline, bitemporal narrowing, low-set protruding ears, strabismus and tented thick eyebrows with sparse hair in their medial segment.

Keywords - Diseaseⁱ

Mental retardation

Organism-specific databases

DisGeNET More...DisGeNETⁱ	55691
MalaCards human disease database More...MalaCardsⁱ	FRMD4A
MIMⁱ	616819 phenotype
Open Targets More...OpenTargetsⁱ	ENSG00000151474
PharmGKBⁱ	PA134946784

Polymorphism and mutation databases

BioMutaⁱ	FRMD4A
DMDMⁱ	205371790

PTM / Processingⁱ

Molecule processing

Feature key	Position(s)	DescriptionActions	Graphical view	Length
ChainⁱPRO_0000219444	1 – 1039	FERM domain-containing protein 4AAdd BLAST		1039

Amino acid modifications

Feature key	Position(s)	DescriptionActions	Length
Modified residueⁱ	530	PhosphoserineCombined sources	1
Modified residueⁱ	604	PhosphoserineCombined sources	1
Modified residueⁱ	615	PhosphoserineCombined sources	1
Modified residueⁱ	681	PhosphoserineCombined sources	1
Modified residueⁱ	711	PhosphoserineBy similarity	1
Modified residueⁱ	800	PhosphoserineCombined sources	1
Modified residueⁱ	872	PhosphoserineCombined sources	1
Modified residueⁱ	901	PhosphoserineCombined sources	1

Keywords - PTMⁱ

Phosphoprotein

Proteomic databases

EPDⁱ	Q9P2Q2
MaxQB - The MaxQuant DataBase More...MaxQBⁱ	Q9P2Q2
PaxDbⁱ	Q9P2Q2
PeptideAtlas More...PeptideAtlasⁱ	Q9P2Q2
PRIDEⁱ	Q9P2Q2
ProteomicsDBⁱ	83879

PTM databases

iPTMnetⁱ	Q9P2Q2
PhosphoSitePlusⁱ	Q9P2Q2

Expressionⁱ

Gene expression databases

Bgeeⁱ	ENSG00000151474
CleanExⁱ	HS_FRMD4A
ExpressionAtlasⁱ	Q9P2Q2 baseline and differential
Genevisibleⁱ	Q9P2Q2 HS

Organism-specific databases

Human Protein Atlas More...HPAⁱ	HPA038449

HPAⁱ

HPA038449

Interactionⁱ

Subunit structureⁱ

Interacts (via coiled-coil domain) with CYTH1 (via coiled-coil domain). Interacts with PARD3 (via coiled-coil domain). Found in a complex with PARD3, CYTH1 and FRMD4A. Interacts with CYTH2. Interacts with CYTH3.By similarity

GO - Molecular functionⁱ

protein binding, bridging Source: Ensembl

View the complete GO annotation on QuickGO ...

Protein-protein interaction databases

BioGridⁱ	120817, 6 interactors
IntActⁱ	Q9P2Q2, 4 interactors
STRINGⁱ	9606.ENSP00000350032

Structureⁱ

3D structure databases

ProteinModelPortalⁱ	Q9P2Q2
SMRⁱ	Q9P2Q2
ModBaseⁱ	Search...
MobiDBⁱ	Search...

Family & Domainsⁱ

Domains and Repeats

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Domainⁱ	20 – 322	FERMPROSITE-ProRule annotationAdd BLAST		303

Region

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Regionⁱ	358 – 420	Necessary for interaction with CYTH1By similarityAdd BLAST		63
Regionⁱ	579 – 939	Necessary for tight junction and adherens junction localization; Requires for interaction with PARD3By similarityAdd BLAST		361

Coiled coil

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Coiled coilⁱ	382 – 416	Sequence analysisAdd BLAST		35

Compositional bias

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Compositional biasⁱ	627 – 756	Ser-richAdd BLAST		130
Compositional biasⁱ	935 – 1007	Ser-richAdd BLAST		73

Keywords - Domainⁱ

Coiled coil

Phylogenomic databases

eggNOGⁱ	KOG3529 Eukaryota ENOG410XQFP LUCA
Ensembl GeneTree More...GeneTreeⁱ	ENSGT00920000149019
HOGENOMⁱ	HOG000294161
HOVERGENⁱ	HBG062800
InParanoidⁱ	Q9P2Q2
OMAⁱ	KEYPLYV
Database of Orthologous Groups More...OrthoDBⁱ	EOG091G0B4Y
PhylomeDBⁱ	Q9P2Q2
TreeFamⁱ	TF328984

Family and domain databases

Conserved Domains Database More...CDDⁱ	cd14473 FERM_B-lobe, 1 hit
Gene3Dⁱ	1.20.80.10, 1 hit 2.30.29.30, 1 hit
InterProⁱ	View protein in InterPro IPR019749 Band_41_domain IPR021774 DUF3338 IPR014352 FERM/acyl-CoA-bd_prot_sf IPR035963 FERM_2 IPR019748 FERM_central IPR019747 FERM_CS IPR000299 FERM_domain IPR018979 FERM_N IPR018980 FERM_PH-like_C IPR011993 PH-like_dom_sf IPR029071 Ubiquitin-like_domsf
Pfam protein domain database More...Pfamⁱ	View protein in Pfam PF11819 DUF3338, 1 hit PF09380 FERM_C, 1 hit PF00373 FERM_M, 1 hit PF09379 FERM_N, 1 hit
PRINTSⁱ	PR00935 BAND41
SMARTⁱ	View protein in SMART SM00295 B41, 1 hit SM01196 FERM_C, 1 hit
SUPFAMⁱ	SSF47031 SSF47031, 1 hit SSF54236 SSF54236, 1 hit
PROSITEⁱ	View protein in PROSITE PS00660 FERM_1, 1 hit PS00661 FERM_2, 1 hit PS50057 FERM_3, 1 hit

Sequenceⁱ

Sequence statusⁱ: Complete.

Q9P2Q2-1 [UniParc]FASTA Add to basket

« Hide

        10         20         30         40         50
MAVQLVPDSA LGLLMMTEGR RCQVHLLDDR KLELLVQPKL LAKELLDLVA 
        60         70         80         90        100
SHFNLKEKEY FGIAFTDETG HLNWLQLDRR VLEHDFPKKS GPVVLYFCVR 
       110        120        130        140        150
FYIESISYLK DNATIELFFL NAKSCIYKEL IDVDSEVVFE LASYILQEAK 
       160        170        180        190        200
GDFSSNEVVR SDLKKLPALP TQALKEHPSL AYCEDRVIEH YKKLNGQTRG 
       210        220        230        240        250
QAIVNYMSIV ESLPTYGVHY YAVKDKQGIP WWLGLSYKGI FQYDYHDKVK 
       260        270        280        290        300
PRKIFQWRQL ENLYFREKKF SVEVHDPRRA SVTRRTFGHS GIAVHTWYAC 
       310        320        330        340        350
PALIKSIWAM AISQHQFYLD RKQSKSKIHA ARSLSEIAID LTETGTLKTS 
       360        370        380        390        400
KLANMGSKGK IISGSSGSLL SSGSQESDSS QSAKKDMLAA LKSRQEALEE 
       410        420        430        440        450
TLRQRLEELK KLCLREAELT GKLPVEYPLD PGEEPPIVRR RIGTAFKLDE 
       460        470        480        490        500
QKILPKGEEA ELERLEREFA IQSQITEAAR RLASDPNVSK KLKKQRKTSY 
       510        520        530        540        550
LNALKKLQEI ENAINENRIK SGKKPTQRAS LIIDDGNIAS EDSSLSDALV 
       560        570        580        590        600
LEDEDSQVTS TISPLHSPHK GLPPRPPSHN RPPPPQSLEG LRQMHYHRND 
       610        620        630        640        650
YDKSPIKPKM WSESSLDEPY EKVKKRSSHS HSSSHKRFPS TGSCAEAGGG 
       660        670        680        690        700
SNSLQNSPIR GLPHWNSQSS MPSTPDLRVR SPHYVHSTRS VDISPTRLHS 
       710        720        730        740        750
LALHFRHRSS SLESQGKLLG SENDTGSPDF YTPRTRSSNG SDPMDDCSSC 
       760        770        780        790        800
TSHSSSEHYY PAQMNANYST LAEDSPSKAR QRQRQRQRAA GALGSASSGS 
       810        820        830        840        850
MPNLAARGGA GGAGGAGGGV YLHSQSQPSS QYRIKEYPLY IEGGATPVVV 
       860        870        880        890        900
RSLESDQEGH YSVKAQFKTS NSYTAGGLFK ESWRGGGGDE GDTGRLTPSR 
       910        920        930        940        950
SQILRTPSLG REGAHDKGAG RAAVSDELRQ WYQRSTASHK EHSRLSHTSS 
       960        970        980        990       1000
TSSDSGSQYS TSSQSTFVAH SRVTRMPQMC KATSAALPQS QRSSTPSSEI 
      1010       1020       1030 
GATPPSSPHH ILTWQTGEAT ENSPILDGSE SPPHQSTDE

Length:1,039

Mass (Da):115,458

Last modified:September 2, 2008 - v3

Checksum:ⁱD895CD6B569A683F

GO

Sequence cautionⁱ

The sequence BAA92532 differs from that shown. Reason: Erroneous initiation.Curated

Experimental Info

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Sequence conflictⁱ	859	G → C in BAA92532 (PubMed:10718198).Curated		1
Sequence conflictⁱ	859	G → C in AAI51245 (PubMed:15489334).Curated		1

Natural variant

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Natural variantⁱVAR_048367	242	Q → H. Corresponds to variant dbSNP:rs11258565Ensembl.		1

Sequence databases

Select the link destinations: EMBLⁱ GenBankⁱ DDBJⁱ Links Updated	AB037715 mRNA Translation: BAA92532.1 Different initiation. AK289693 mRNA Translation: BAF82382.1 AC069025 Genomic DNA No translation available. AL157392 Genomic DNA No translation available. AL365495 Genomic DNA No translation available. BC151244 mRNA Translation: AAI51245.1
The Consensus CDS (CCDS) project More...CCDSⁱ	CCDS7101.1
NCBI Reference Sequences More...RefSeqⁱ	NP_001305265.1, NM_001318336.1 NP_001305266.1, NM_001318337.1 NP_001305267.1, NM_001318338.1 NP_060497.3, NM_018027.4
UniGeneⁱ	Hs.330463 Hs.568818 Hs.656573

Genome annotation databases

Ensemblⁱ	ENST00000357447; ENSP00000350032; ENSG00000151474
GeneIDⁱ	55691
KEGGⁱ	hsa:55691
UCSC genome browser More...UCSCⁱ	uc001ims.4 human

Keywords - Coding sequence diversityⁱ

Polymorphism

Similar proteinsⁱ

90% Identity
50% Identity

Protein	Similar proteins		Species	Score	Length
Q9P2Q2	UPI0007B41C0B		MONDO		1113
FERM domain-containing protein 4A isoform X2		AMAAE		1091
UPI000462C802		ANOCA		1090
FERM domain-containing protein 4A		ODORO		1087
UPI000670B571		Anser cygnoides domesticus		1084
+394

Protein	Similar proteins		Species	Score	Length
Q9P2Q2	UPI0007B41C0B		MONDO		1113
FERM domain-containing protein 4A isoform X2		AMAAE		1091
UPI000462C802		ANOCA		1090
FERM domain-containing protein 4A		ODORO		1087
UPI000670B571		Anser cygnoides domesticus		1084
+636

Cross-referencesⁱ

Sequence databases

Select the link destinations: EMBLⁱ GenBankⁱ DDBJⁱ Links Updated	AB037715 mRNA Translation: BAA92532.1 Different initiation. AK289693 mRNA Translation: BAF82382.1 AC069025 Genomic DNA No translation available. AL157392 Genomic DNA No translation available. AL365495 Genomic DNA No translation available. BC151244 mRNA Translation: AAI51245.1
The Consensus CDS (CCDS) project More...CCDSⁱ	CCDS7101.1
NCBI Reference Sequences More...RefSeqⁱ	NP_001305265.1, NM_001318336.1 NP_001305266.1, NM_001318337.1 NP_001305267.1, NM_001318338.1 NP_060497.3, NM_018027.4
UniGeneⁱ	Hs.330463 Hs.568818 Hs.656573

3D structure databases

ProteinModelPortalⁱ	Q9P2Q2
SMRⁱ	Q9P2Q2
ModBaseⁱ	Search...
MobiDBⁱ	Search...

Protein-protein interaction databases

BioGridⁱ	120817, 6 interactors
IntActⁱ	Q9P2Q2, 4 interactors
STRINGⁱ	9606.ENSP00000350032

PTM databases

iPTMnetⁱ	Q9P2Q2
PhosphoSitePlusⁱ	Q9P2Q2

Polymorphism and mutation databases

BioMutaⁱ	FRMD4A
DMDMⁱ	205371790

Proteomic databases

EPDⁱ	Q9P2Q2
MaxQB - The MaxQuant DataBase More...MaxQBⁱ	Q9P2Q2
PaxDbⁱ	Q9P2Q2
PeptideAtlas More...PeptideAtlasⁱ	Q9P2Q2
PRIDEⁱ	Q9P2Q2
ProteomicsDBⁱ	83879

Protocols and materials databases

Structural Biology Knowledgebase	Search...

Structural Biology Knowledgebase

Search...

Genome annotation databases

Ensemblⁱ	ENST00000357447; ENSP00000350032; ENSG00000151474
GeneIDⁱ	55691
KEGGⁱ	hsa:55691
UCSC genome browser More...UCSCⁱ	uc001ims.4 human

Organism-specific databases

CTDⁱ	55691
DisGeNET More...DisGeNETⁱ	55691
EuPathDBⁱ	HostDB:ENSG00000151474.19
GeneCardsⁱ	FRMD4A
H-InvDBⁱ	HIX0008660
Human Gene Nomenclature Database More...HGNCⁱ	HGNC:25491 FRMD4A
Human Protein Atlas More...HPAⁱ	HPA038449
MalaCards human disease database More...MalaCardsⁱ	FRMD4A
MIMⁱ	616305 gene 616819 phenotype
neXtProtⁱ	NX_Q9P2Q2
Open Targets More...OpenTargetsⁱ	ENSG00000151474
PharmGKBⁱ	PA134946784
HUGEⁱ	Search...
GenAtlas: human gene database More...GenAtlasⁱ	Search...

Phylogenomic databases

eggNOGⁱ	KOG3529 Eukaryota ENOG410XQFP LUCA
Ensembl GeneTree More...GeneTreeⁱ	ENSGT00920000149019
HOGENOMⁱ	HOG000294161
HOVERGENⁱ	HBG062800
InParanoidⁱ	Q9P2Q2
OMAⁱ	KEYPLYV
Database of Orthologous Groups More...OrthoDBⁱ	EOG091G0B4Y
PhylomeDBⁱ	Q9P2Q2
TreeFamⁱ	TF328984

Miscellaneous databases

ChiTaRSⁱ	FRMD4A human
GenomeRNAiⁱ	55691
Protein Ontology More...PROⁱ	PR:Q9P2Q2
SOURCEⁱ	Search...

Gene expression databases

Bgeeⁱ	ENSG00000151474
CleanExⁱ	HS_FRMD4A
ExpressionAtlasⁱ	Q9P2Q2 baseline and differential
Genevisibleⁱ	Q9P2Q2 HS

Family and domain databases

Conserved Domains Database More...CDDⁱ	cd14473 FERM_B-lobe, 1 hit
Gene3Dⁱ	1.20.80.10, 1 hit 2.30.29.30, 1 hit
InterProⁱ	View protein in InterPro IPR019749 Band_41_domain IPR021774 DUF3338 IPR014352 FERM/acyl-CoA-bd_prot_sf IPR035963 FERM_2 IPR019748 FERM_central IPR019747 FERM_CS IPR000299 FERM_domain IPR018979 FERM_N IPR018980 FERM_PH-like_C IPR011993 PH-like_dom_sf IPR029071 Ubiquitin-like_domsf
Pfam protein domain database More...Pfamⁱ	View protein in Pfam PF11819 DUF3338, 1 hit PF09380 FERM_C, 1 hit PF00373 FERM_M, 1 hit PF09379 FERM_N, 1 hit
PRINTSⁱ	PR00935 BAND41
SMARTⁱ	View protein in SMART SM00295 B41, 1 hit SM01196 FERM_C, 1 hit
SUPFAMⁱ	SSF47031 SSF47031, 1 hit SSF54236 SSF54236, 1 hit
PROSITEⁱ	View protein in PROSITE PS00660 FERM_1, 1 hit PS00661 FERM_2, 1 hit PS50057 FERM_3, 1 hit
ProtoNetⁱ	Search...

Entry informationⁱ

Entry nameⁱ	FRM4A_HUMAN
Accessionⁱ	Q9P2Q2Primary (citable) accession number: Q9P2Q2 Secondary accession number(s): A7E2Y3, Q5T377
Entry historyⁱ	Integrated into UniProtKB/Swiss-Prot:	July 11, 2001
	Last sequence update:	September 2, 2008
	Last modified:	June 20, 2018
	This is version 138 of the entry and version 3 of the sequence. See complete history.
Entry statusⁱ	Reviewed (UniProtKB/Swiss-Prot)
Annotation program	Chordata Protein Annotation Program
Disclaimer	Any medical or genetic information present in this entry is provided for research, educational and informational purposes only. It is not in any way intended to be used as a substitute for professional medical advice, diagnosis, treatment or care.

Miscellaneousⁱ

Keywords - Technical termⁱ

Complete proteome, Reference proteome

Documents

Human chromosome 10
Human chromosome 10: entries, gene names and cross-references to MIM
Human entries with polymorphisms or disease mutations
List of human entries with polymorphisms or disease mutations
Human polymorphisms and disease mutations
Index of human polymorphisms and disease mutations
MIM cross-references
Online Mendelian Inheritance in Man (MIM) cross-references in UniProtKB/Swiss-Prot

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Supporting data

UniProtKB - Q9P2Q2 (FRM4A_HUMAN)

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FERM domain-containing protein 4A

Select a section on the left to see content.

<p>This section provides any useful information about the protein, mostly biological knowledge.<p><a href='/help/function_section' target='_top'>More...</a></p>Functioni

Caution

<p>The <a href="http://www.geneontology.org/">Gene Ontology (GO)</a> project provides a set of hierarchical controlled vocabulary split into 3 categories:<p><a href='/help/gene_ontology' target='_top'>More...</a></p>GO - Molecular functioni

<p>The <a href="http://www.geneontology.org/">Gene Ontology (GO)</a> project provides a set of hierarchical controlled vocabulary split into 3 categories:<p><a href='/help/gene_ontology' target='_top'>More...</a></p>GO - Biological processi

<p>This section provides information about the protein and gene name(s) and synonym(s) and about the organism that is the source of the protein sequence.<p><a href='/help/names_and_taxonomy_section' target='_top'>More...</a></p>Names & Taxonomyi

Organism-specific databases

<p>This section provides information on the location and the topology of the mature protein in the cell.<p><a href='/help/subcellular_location_section' target='_top'>More...</a></p>Subcellular locationi

Cytoskeleton

Other locations

Cytoskeleton

Other locations

<p>This section provides information on the disease(s) and phenotype(s) associated with a protein.<p><a href='/help/pathology_and_biotech_section' target='_top'>More...</a></p>Pathology & Biotechi

Organism-specific databases

Polymorphism and mutation databases

<p>This section describes post-translational modifications (PTMs) and/or processing events.<p><a href='/help/ptm_processing_section' target='_top'>More...</a></p>PTM / Processingi

Molecule processing

Amino acid modifications

Proteomic databases

PTM databases

<p>This section provides information on the expression of a gene at the mRNA or protein level in cells or in tissues of multicellular organisms.<p><a href='/help/expression_section' target='_top'>More...</a></p>Expressioni

Gene expression databases

Organism-specific databases

<p>This section provides information on the quaternary structure of a protein and on interaction(s) with other proteins or protein complexes.<p><a href='/help/interaction_section' target='_top'>More...</a></p>Interactioni

<p>The <a href="http://www.geneontology.org/">Gene Ontology (GO)</a> project provides a set of hierarchical controlled vocabulary split into 3 categories:<p><a href='/help/gene_ontology' target='_top'>More...</a></p>GO - Molecular functioni

Protein-protein interaction databases

<p>This section provides information on the tertiary and secondary structure of a protein.<p><a href='/help/structure_section' target='_top'>More...</a></p>Structurei

3D structure databases

<p>This section provides information on sequence similarities with other proteins and the domain(s) present in a protein.<p><a href='/help/family_and_domains_section' target='_top'>More...</a></p>Family & Domainsi

Domains and Repeats

Region

Coiled coil

Compositional bias

Phylogenomic databases

Family and domain databases

<p>This subsection of the ‘Sequence’ section reports difference(s) between the protein sequence shown in the UniProtKB entry and other available protein sequences derived from the same gene.<p><a href='/help/sequence_caution' target='_top'>More...</a></p>Sequence cautioni

Experimental Info

Natural variant

Sequence databases

Genome annotation databases

<p>This section is used to point to information related to entries and found in data collections other than UniProtKB.<p><a href='/help/cross_references_section' target='_top'>More...</a></p>Cross-referencesi

Sequence databases

3D structure databases

Protein-protein interaction databases

PTM databases

Polymorphism and mutation databases

Proteomic databases

Protocols and materials databases

Genome annotation databases

Organism-specific databases

Phylogenomic databases

Miscellaneous databases

Gene expression databases

Family and domain databases

<p>This section provides general information on the entry.<p><a href='/help/entry_information_section' target='_top'>More...</a></p>Entry informationi

<p>This section contains any relevant information that doesn’t fit in any other defined sections<p><a href='/help/miscellaneous_section' target='_top'>More...</a></p>Miscellaneousi

Documents

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Functionⁱ

GO - Molecular functionⁱ

GO - Biological processⁱ

Names & Taxonomyⁱ

Subcellular locationⁱ

Pathology & Biotechⁱ

PTM / Processingⁱ

Expressionⁱ

Interactionⁱ

GO - Molecular functionⁱ

Structureⁱ

Family & Domainsⁱ

Sequence cautionⁱ

Cross-referencesⁱ

Entry informationⁱ

Miscellaneousⁱ