MAL

Functionⁱ

Could be an important component in vesicular trafficking cycling between the Golgi complex and the apical plasma membrane. Could be involved in myelin biogenesis and/or myelin function.

GO - Molecular functionⁱ

lipid binding
Source: UniProtKB
Traceable author statementⁱ
- 10739088
peptidase activator activity involved in apoptotic process
Source: UniProtKB
Non-traceable author statementⁱ
- 10366425
structural constituent of myelin sheath
Source: UniProtKB
Inferred from direct assayⁱ
- 12153479

View the complete GO annotation on QuickGO ...

GO - Biological processⁱ

apical protein localization
Source: UniProtKB
Traceable author statementⁱ
- 10739088
apoptotic process
Source: UniProtKB
Non-traceable author statementⁱ
- 10366425
cell differentiation
Source: UniProtKB
Traceable author statementⁱ
- 10739088
central nervous system development
Source: UniProtKB
Traceable author statementⁱ
- 10739088
membrane raft polarization
Source: UniProtKB
Traceable author statementⁱ
- 12153479
myelination
Source: GO_Central
Inferred from biological aspect of ancestorⁱ
- 21873635
positive regulation of extrinsic apoptotic signaling pathway via death domain receptors Source: Ensembl
protein insertion into plasma membrane Source: Ensembl
protein localization to paranode region of axon Source: Ensembl

View the complete GO annotation on QuickGO ...

Names & Taxonomyⁱ

Protein namesⁱ	Recommended name: Myelin and lymphocyte protein Alternative name(s): T-lymphocyte maturation-associated protein
Gene namesⁱ	Name:MAL
Organismⁱ	Homo sapiens (Human)
Taxonomic identifierⁱ	9606 [NCBI]
Taxonomic lineageⁱ	cellular organisms › Eukaryota › Opisthokonta › Metazoa › Eumetazoa › Bilateria › Deuterostomia › Chordata › Craniata › Vertebrata › Gnathostomata › Teleostomi › Euteleostomi › Sarcopterygii › Dipnotetrapodomorpha › Tetrapoda › Amniota › Mammalia › Theria › Eutheria › Boreoeutheria › Euarchontoglires › Primates › Haplorrhini › Simiiformes › Catarrhini › Hominoidea › Hominidae › Homininae › Homo
Proteomesⁱ	UP000005640 Componentⁱ: Chromosome 2

Organism-specific databases

EuPathDBⁱ	HostDB:ENSG00000172005.10
Human Gene Nomenclature Database More...HGNCⁱ	HGNC:6817 MAL
MIMⁱ	188860 gene
neXtProtⁱ	NX_P21145

Topology

Feature key	Position(s)	DescriptionActions	Length
Topological domainⁱ	1 – 24	CytoplasmicSequence analysisAdd BLAST	24
Transmembraneⁱ	25 – 46	HelicalSequence analysisAdd BLAST	22
Topological domainⁱ	47 – 53	ExtracellularSequence analysis	7
Transmembraneⁱ	54 – 75	HelicalSequence analysisAdd BLAST	22
Topological domainⁱ	76 – 92	CytoplasmicSequence analysisAdd BLAST	17
Transmembraneⁱ	93 – 114	HelicalSequence analysisAdd BLAST	22
Topological domainⁱ	115 – 125	ExtracellularSequence analysisAdd BLAST	11
Transmembraneⁱ	126 – 147	HelicalSequence analysisAdd BLAST	22
Topological domainⁱ	148 – 153	CytoplasmicSequence analysis	6

Keywords - Cellular componentⁱ

Membrane

Pathology & Biotechⁱ

Organism-specific databases

DisGeNET More...DisGeNETⁱ	4118
Open Targets More...OpenTargetsⁱ	ENSG00000172005
PharmGKBⁱ	PA30565

Polymorphism and mutation databases

BioMutaⁱ	MAL
DMDMⁱ	126719

PTM / Processingⁱ

Molecule processing

Feature key	Position(s)	DescriptionActions	Graphical view	Length
ChainⁱPRO_0000156805	1 – 153	Myelin and lymphocyte proteinAdd BLAST		153

Post-translational modificationⁱ

Lipoprotein.

Keywords - PTMⁱ

Lipoprotein

Proteomic databases

PaxDbⁱ	P21145
PeptideAtlas More...PeptideAtlasⁱ	P21145
PRIDEⁱ	P21145
ProteomicsDBⁱ	53849 53850 [P21145-2] 53851 [P21145-3] 53852 [P21145-4]

Expressionⁱ

Developmental stageⁱ

Expressed in the intermediate and late stages of T-cell differentiation.

Gene expression databases

Bgeeⁱ	ENSG00000172005 Expressed in 207 organ(s), highest expression level in esophagus squamous epithelium
CleanExⁱ	HS_MAL
ExpressionAtlasⁱ	P21145 baseline and differential
Genevisibleⁱ	P21145 HS

Organism-specific databases

Human Protein Atlas More...HPAⁱ	HPA065691

HPAⁱ

HPA065691

Interactionⁱ

Binary interactionsⁱ

With	Entry	#Exp.	IntAct
BTK	Q06187	5	EBI-3932027,EBI-624835
EDA	Q92838	5	EBI-3932027,EBI-529425
FIBCD1	Q8N539	5	EBI-3932027,EBI-10200808
KLRC1	P26715	5	EBI-3932027,EBI-9018187
LDLRAD1	Q5T700	5	EBI-3932027,EBI-10173166
LSMEM1	Q8N8F7	3	EBI-3932027,EBI-10200825
PTPRN	Q16849-3	3	EBI-3932027,EBI-10200782
SMIM3	Q9BZL3	5	EBI-3932027,EBI-741850
USP7	Q93009	3	EBI-3932027,EBI-302474

Protein-protein interaction databases

BioGridⁱ	110292, 10 interactors
IntActⁱ	P21145, 74 interactors
STRINGⁱ	9606.ENSP00000310880

Structureⁱ

3D structure databases

ProteinModelPortalⁱ	P21145
ModBaseⁱ	Search...
MobiDBⁱ	Search...

Family & Domainsⁱ

Domains and Repeats

Feature key	Position(s)	DescriptionActions	Graphical view	Length
Domainⁱ	18 – 151	MARVELPROSITE-ProRule annotationAdd BLAST		134

Sequence similaritiesⁱ

Belongs to the MAL family.Curated

Keywords - Domainⁱ

Transmembrane, Transmembrane helix

Phylogenomic databases

eggNOGⁱ	KOG4788 Eukaryota ENOG4111IAF LUCA
Ensembl GeneTree More...GeneTreeⁱ	ENSGT00730000110762
HOGENOMⁱ	HOG000231752
HOVERGENⁱ	HBG000925
InParanoidⁱ	P21145
OMAⁱ	LIQGWVM
Database of Orthologous Groups More...OrthoDBⁱ	EOG091G0OS1
PhylomeDBⁱ	P21145
TreeFamⁱ	TF316174

Family and domain databases

InterProⁱ	View protein in InterPro IPR013295 MAL IPR008253 Marvel
Pfam protein domain database More...Pfamⁱ	View protein in Pfam PF01284 MARVEL, 1 hit
PRINTSⁱ	PR01884 MALPROTEIN
PROSITEⁱ	View protein in PROSITE PS51225 MARVEL, 1 hit

Sequences (4)ⁱ

Sequence statusⁱ: Complete.

This entry describes 4 isoformsⁱ produced by alternative splicing. Align Add to basket

Isoform A (identifier: P21145-1) [UniParc]FASTA Add to basket

This isoform has been chosen as the 'canonical' sequence. All positional information in this entry refers to it. This is also the sequence that appears in the downloadable versions of the entry.

« Hide

        10         20         30         40         50
MAPAAATGGS TLPSGFSVFT TLPDLLFIFE FIFGGLVWIL VASSLVPWPL 
        60         70         80         90        100
VQGWVMFVSV FCFVATTTLI ILYIIGAHGG ETSWVTLDAA YHCTAALFYL 
       110        120        130        140        150
SASVLEALAT ITMQDGFTYR HYHENIAAVV FSYIATLLYV VHAVFSLIRW 

KSS

Length:153

Mass (Da):16,714

Last modified:May 1, 1991 - v1

Checksum:ⁱF821FC1C0ADBE775

GO

Isoform B (identifier: P21145-2) [UniParc]FASTA Add to basket

The sequence of this isoform differs from the canonical sequence as follows:
88-129: Missing.

« Hide

        10         20         30         40         50
MAPAAATGGS TLPSGFSVFT TLPDLLFIFE FIFGGLVWIL VASSLVPWPL 
        60         70         80         90        100
VQGWVMFVSV FCFVATTTLI ILYIIGAHGG ETSWVTLVFS YIATLLYVVH 
       110 
AVFSLIRWKS S

Show »

Length:111

Mass (Da):12,081

Checksum:ⁱ076826F85AF6E58B

GO

Isoform C (identifier: P21145-3) [UniParc]FASTA Add to basket

The sequence of this isoform differs from the canonical sequence as follows:
32-87: Missing.

« Hide

        10         20         30         40         50
MAPAAATGGS TLPSGFSVFT TLPDLLFIFE FDAAYHCTAA LFYLSASVLE 
        60         70         80         90 
ALATITMQDG FTYRHYHENI AAVVFSYIAT LLYVVHAVFS LIRWKSS

Show »

Length:97

Mass (Da):10,621

Checksum:ⁱ831D98D55230C2DC

GO

Isoform D (identifier: P21145-4) [UniParc]FASTA Add to basket

The sequence of this isoform differs from the canonical sequence as follows:
32-129: Missing.

« Hide

        10         20         30         40         50
MAPAAATGGS TLPSGFSVFT TLPDLLFIFE FVFSYIATLL YVVHAVFSLI 

RWKSS

Show »

Length:55

Mass (Da):5,989

Checksum:ⁱF943BC95A3AF4927

GO

Alternative sequence

Feature key	Position(s)	DescriptionActions	Length
Alternative sequenceⁱVSP_003162	32 – 129	Missing in isoform D. CuratedAdd BLAST	98
Alternative sequenceⁱVSP_003163	32 – 87	Missing in isoform C. CuratedAdd BLAST	56
Alternative sequenceⁱVSP_003164	88 – 129	Missing in isoform B. CuratedAdd BLAST	42

Sequence databases

Select the link destinations: EMBLⁱ GenBankⁱ DDBJⁱ Links Updated	M15800 mRNA Translation: AAA36196.1 X76678 mRNA Translation: CAA54100.1 X76679 mRNA Translation: CAA54101.1 X76680 mRNA Translation: CAA54102.1 X76681 mRNA Translation: CAA54103.1 X76220 , X76221, X76222, X76223 Genomic DNA Translation: CAA53809.1 AK311844 mRNA Translation: BAG34786.1 CR541879 mRNA Translation: CAG46677.1 AC103563 Genomic DNA Translation: AAY24121.1 CH471219 Genomic DNA Translation: EAX10705.1 BC000458 mRNA Translation: AAH00458.1 BC003006 mRNA Translation: AAH03006.1
The Consensus CDS (CCDS) project More...CCDSⁱ	CCDS2006.1 [P21145-1] CCDS2007.1 [P21145-2] CCDS2008.1 [P21145-3] CCDS2009.1 [P21145-4]
PIRⁱ	A29472
NCBI Reference Sequences More...RefSeqⁱ	NP_002362.1, NM_002371.3 [P21145-1] NP_071883.1, NM_022438.2 [P21145-2] NP_071884.1, NM_022439.2 [P21145-3] NP_071885.1, NM_022440.2 [P21145-4]
UniGeneⁱ	Hs.80395

Genome annotation databases

Ensemblⁱ	ENST00000309988; ENSP00000310880; ENSG00000172005 [P21145-1] ENST00000349807; ENSP00000322860; ENSG00000172005 [P21145-4] ENST00000353004; ENSP00000306568; ENSG00000172005 [P21145-2] ENST00000354078; ENSP00000304924; ENSG00000172005 [P21145-3]
GeneIDⁱ	4118
KEGGⁱ	hsa:4118
UCSC genome browser More...UCSCⁱ	uc002stx.3 human [P21145-1]

Keywords - Coding sequence diversityⁱ

Alternative splicing

Similar proteinsⁱ

100% Identity
90% Identity
50% Identity

Protein	Similar proteins		Species	Score	Length	Source
P21145	Mal, T-cell differentiation protein, isoform CRA_a		HUMAN		153	UniRef100_P21145

Protein	Similar proteins		Species	Score	Length	Source
P21145	Mal, T-cell differentiation protein, isoform CRA_a		HUMAN		153	UniRef90_P21145
MAL isoform 2		PANTR		153
Uncharacterized protein		PANPA		153
Mal, T cell differentiation protein		GORGO		153
Uncharacterized protein		MACMU		153
+17
P21145-2	MAL isoform 5		PANTR		111	UniRef90_P21145-2
Mal, T cell differentiation protein		GORGO		111
Uncharacterized protein		CALJA		111
Mal, T cell differentiation protein		SAIBB		111
Uncharacterized protein		PROCO		111
+15
P21145-3	MAL isoform 4		PANTR		97	UniRef90_P21145-3
Mal, T cell differentiation protein		GORGO		97
Mal, T cell differentiation protein		SAIBB		97
Uncharacterized protein		CALJA		97
Uncharacterized protein		MACMU		97
+13
P21145-4	MAL isoform 3		PANTR		55	UniRef90_P21145-4
Mal, T cell differentiation protein		GORGO		55
Uncharacterized protein		CALJA		55
Mal, T cell differentiation protein		SAIBB		55
Uncharacterized protein		MACMU		55
+21

Protein	Similar proteins		Species	Score	Length	Source
P21145	Myelin and lymphocyte protein		CANLF		153	UniRef50_P21145
Myelin and lymphocyte protein		RAT		153
Myelin and lymphocyte protein		MOUSE		153
Myelin and lymphocyte protein		BOVIN		153
Mal, T-cell differentiation protein, isoform CRA_a		HUMAN		153
+180
P21145-2	Uncharacterized protein		CALJA		111	UniRef50_P21145-2
Mal, T cell differentiation protein		SAIBB		111
MAL isoform 5		PONAB		111
myelin and lymphocyte protein isoform X2		ODORO		111
Uncharacterized protein		Cebus capucinus imitator		111
+55
P21145-3	Uncharacterized protein		CALJA		97	UniRef50_P21145-3
Uncharacterized protein		NOMLE		97
Uncharacterized protein		Cebus capucinus imitator		97
MAL isoform 4		PANTR		97
Mal, T cell differentiation protein		GORGO		97
+33
P21145-4	myelin and lymphocyte protein isoform X4		NEOSC		55	UniRef50_P21145-4
Uncharacterized protein		AOTNA		55
Uncharacterized protein		FELCA		55
myelin and lymphocyte protein isoform X4		ODORO		55
MAL isoform 3		PANTR		55
+48

Cross-referencesⁱ

Web resourcesⁱ

Atlas of Genetics and Cytogenetics in Oncology and Haematology

Sequence databases

Select the link destinations: EMBLⁱ GenBankⁱ DDBJⁱ Links Updated	M15800 mRNA Translation: AAA36196.1 X76678 mRNA Translation: CAA54100.1 X76679 mRNA Translation: CAA54101.1 X76680 mRNA Translation: CAA54102.1 X76681 mRNA Translation: CAA54103.1 X76220 , X76221, X76222, X76223 Genomic DNA Translation: CAA53809.1 AK311844 mRNA Translation: BAG34786.1 CR541879 mRNA Translation: CAG46677.1 AC103563 Genomic DNA Translation: AAY24121.1 CH471219 Genomic DNA Translation: EAX10705.1 BC000458 mRNA Translation: AAH00458.1 BC003006 mRNA Translation: AAH03006.1
CCDSⁱ	CCDS2006.1 [P21145-1] CCDS2007.1 [P21145-2] CCDS2008.1 [P21145-3] CCDS2009.1 [P21145-4]
PIRⁱ	A29472
RefSeqⁱ	NP_002362.1, NM_002371.3 [P21145-1] NP_071883.1, NM_022438.2 [P21145-2] NP_071884.1, NM_022439.2 [P21145-3] NP_071885.1, NM_022440.2 [P21145-4]
UniGeneⁱ	Hs.80395

3D structure databases

ProteinModelPortalⁱ	P21145
ModBaseⁱ	Search...
MobiDBⁱ	Search...

Protein-protein interaction databases

BioGridⁱ	110292, 10 interactors
IntActⁱ	P21145, 74 interactors
STRINGⁱ	9606.ENSP00000310880

Polymorphism and mutation databases

BioMutaⁱ	MAL
DMDMⁱ	126719

Proteomic databases

PaxDbⁱ	P21145
PeptideAtlasⁱ	P21145
PRIDEⁱ	P21145
ProteomicsDBⁱ	53849 53850 [P21145-2] 53851 [P21145-3] 53852 [P21145-4]

Protocols and materials databases

Structural Biology Knowledgebase	Search...

Structural Biology Knowledgebase

Search...

Genome annotation databases

Ensemblⁱ	ENST00000309988; ENSP00000310880; ENSG00000172005 [P21145-1] ENST00000349807; ENSP00000322860; ENSG00000172005 [P21145-4] ENST00000353004; ENSP00000306568; ENSG00000172005 [P21145-2] ENST00000354078; ENSP00000304924; ENSG00000172005 [P21145-3]
GeneIDⁱ	4118
KEGGⁱ	hsa:4118
UCSCⁱ	uc002stx.3 human [P21145-1]

Organism-specific databases

CTDⁱ	4118
DisGeNETⁱ	4118
EuPathDBⁱ	HostDB:ENSG00000172005.10
GeneCardsⁱ	MAL
H-InvDBⁱ	HIX0117680 HIX0117687
HGNCⁱ	HGNC:6817 MAL
HPAⁱ	HPA065691
MIMⁱ	188860 gene
neXtProtⁱ	NX_P21145
OpenTargetsⁱ	ENSG00000172005
PharmGKBⁱ	PA30565
GenAtlas: human gene database More...GenAtlasⁱ	Search...

Phylogenomic databases

eggNOGⁱ	KOG4788 Eukaryota ENOG4111IAF LUCA
GeneTreeⁱ	ENSGT00730000110762
HOGENOMⁱ	HOG000231752
HOVERGENⁱ	HBG000925
InParanoidⁱ	P21145
OMAⁱ	LIQGWVM
OrthoDBⁱ	EOG091G0OS1
PhylomeDBⁱ	P21145
TreeFamⁱ	TF316174

Miscellaneous databases

ChiTaRSⁱ	MAL human
GeneWikiⁱ	MAL_(gene)
GenomeRNAiⁱ	4118
Protein Ontology More...PROⁱ	PR:P21145
SOURCEⁱ	Search...

Gene expression databases

Bgeeⁱ	ENSG00000172005 Expressed in 207 organ(s), highest expression level in esophagus squamous epithelium
CleanExⁱ	HS_MAL
ExpressionAtlasⁱ	P21145 baseline and differential
Genevisibleⁱ	P21145 HS

Family and domain databases

InterProⁱ	View protein in InterPro IPR013295 MAL IPR008253 Marvel
Pfamⁱ	View protein in Pfam PF01284 MARVEL, 1 hit
PRINTSⁱ	PR01884 MALPROTEIN
PROSITEⁱ	View protein in PROSITE PS51225 MARVEL, 1 hit
ProtoNetⁱ	Search...

Entry informationⁱ

Entry nameⁱ	MAL_HUMAN
Accessionⁱ	P21145Primary (citable) accession number: P21145 Secondary accession number(s): Q6FH77
Entry historyⁱ	Integrated into UniProtKB/Swiss-Prot:	May 1, 1991
	Last sequence update:	May 1, 1991
	Last modified:	November 7, 2018
	This is version 160 of the entry and version 1 of the sequence. See complete history.
Entry statusⁱ	Reviewed (UniProtKB/Swiss-Prot)
Annotation program	Chordata Protein Annotation Program
Disclaimer	Any medical or genetic information present in this entry is provided for research, educational and informational purposes only. It is not in any way intended to be used as a substitute for professional medical advice, diagnosis, treatment or care.

Miscellaneousⁱ

Keywords - Technical termⁱ

Complete proteome, Reference proteome

Documents

SIMILARITY comments
Index of protein domains and families
Human chromosome 2
Human chromosome 2: entries, gene names and cross-references to MIM
MIM cross-references
Online Mendelian Inheritance in Man (MIM) cross-references in UniProtKB/Swiss-Prot

UniProtKB

UniParc

Help

UniRef

Proteomes

Annotation systems

Supporting data

UniProtKB - P21145 (MAL_HUMAN)

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Myelin and lymphocyte protein

Select a section on the left to see content.

<p>This section provides any useful information about the protein, mostly biological knowledge.<p><a href='/help/function_section' target='_top'>More...</a></p>Functioni

<p>The <a href="http://www.geneontology.org/">Gene Ontology (GO)</a> project provides a set of hierarchical controlled vocabulary split into 3 categories:<p><a href='/help/gene_ontology' target='_top'>More...</a></p>GO - Molecular functioni

GO - Biological processi

<p>This section provides information about the protein and gene name(s) and synonym(s) and about the organism that is the source of the protein sequence.<p><a href='/help/names_and_taxonomy_section' target='_top'>More...</a></p>Names & Taxonomyi

Organism-specific databases

<p>This section provides information on the location and the topology of the mature protein in the cell.<p><a href='/help/subcellular_location_section' target='_top'>More...</a></p>Subcellular locationi

Other locations

Endoplasmic reticulum

Plasma Membrane

Other locations

Topology

<p>This section provides information on the disease(s) and phenotype(s) associated with a protein.<p><a href='/help/pathology_and_biotech_section' target='_top'>More...</a></p>Pathology & Biotechi

Organism-specific databases

Polymorphism and mutation databases

<p>This section describes post-translational modifications (PTMs) and/or processing events.<p><a href='/help/ptm_processing_section' target='_top'>More...</a></p>PTM / Processingi

Molecule processing

Keywords - PTMi

Proteomic databases

<p>This section provides information on the expression of a gene at the mRNA or protein level in cells or in tissues of multicellular organisms.<p><a href='/help/expression_section' target='_top'>More...</a></p>Expressioni

Gene expression databases

Organism-specific databases

<p>This section provides information on the quaternary structure of a protein and on interaction(s) with other proteins or protein complexes.<p><a href='/help/interaction_section' target='_top'>More...</a></p>Interactioni

Protein-protein interaction databases

<p>This section provides information on the tertiary and secondary structure of a protein.<p><a href='/help/structure_section' target='_top'>More...</a></p>Structurei

3D structure databases

<p>This section provides information on sequence similarities with other proteins and the domain(s) present in a protein.<p><a href='/help/family_and_domains_section' target='_top'>More...</a></p>Family & Domainsi

Domains and Repeats

<p>This subsection of the ‘Family and domains’ section provides information about the sequence similarity with other proteins.<p><a href='/help/sequence_similarities' target='_top'>More...</a></p>Sequence similaritiesi

Keywords - Domaini

Phylogenomic databases

Family and domain databases

Alternative sequence

Sequence databases

Genome annotation databases

Keywords - Coding sequence diversityi

<p>This section is used to point to information related to entries and found in data collections other than UniProtKB.<p><a href='/help/cross_references_section' target='_top'>More...</a></p>Cross-referencesi

<p>This subsection of the <a href="http://www.uniprot.org/manual/cross_references_section">Cross-references</a> section provides links to various web resources that are relevant for a specific protein.<p><a href='/help/web_resource' target='_top'>More...</a></p>Web resourcesi

Sequence databases

3D structure databases

Protein-protein interaction databases

Polymorphism and mutation databases

Proteomic databases

Protocols and materials databases

Genome annotation databases

Organism-specific databases

Phylogenomic databases

Miscellaneous databases

Gene expression databases

Family and domain databases

<p>This section provides general information on the entry.<p><a href='/help/entry_information_section' target='_top'>More...</a></p>Entry informationi

<p>This section contains any relevant information that doesn’t fit in any other defined sections<p><a href='/help/miscellaneous_section' target='_top'>More...</a></p>Miscellaneousi

Keywords - Technical termi

Documents

UniProt is an ELIXIR core data resource

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Functionⁱ

GO - Molecular functionⁱ

GO - Biological processⁱ

Names & Taxonomyⁱ

Subcellular locationⁱ

Pathology & Biotechⁱ

PTM / Processingⁱ

Keywords - PTMⁱ

Expressionⁱ

Interactionⁱ

Structureⁱ

Family & Domainsⁱ

Sequence similaritiesⁱ

Keywords - Domainⁱ

Keywords - Coding sequence diversityⁱ

Cross-referencesⁱ

Web resourcesⁱ

Entry informationⁱ

Miscellaneousⁱ

Keywords - Technical termⁱ