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StatusReference proteome
Proteinsi <p>Number of protein entries associated with this proteome: UniProtKB entries for regular proteomes or UniParc entries for redundant proteomes (<a href="/help/proteome%5Fredundancy">more...</a>)</p> 30,858
Gene counti <p>This is the total number of unique genes found in the proteome set, algorithmically computed. For each gene, a single representative protein sequence is chosen from the proteome. Where possible, reviewed (Swiss-Prot) protein sequences are chosen as the representatives.</p> - Download one protein sequence per gene (FASTA)
Proteome IDi <p>The proteome identifier (UPID) is the unique identifier assigned to the set of proteins that constitute the <a href="">proteome</a>. It consists of the characters 'UP' followed by 9 digits, is stable across releases and can therefore be used to cite a UniProt proteome.<p><a href='/help/proteome_id' target='_top'>More...</a></p>UP000006727
Taxonomy3218 - Physcomitrella patens subsp. patens
Straincv. Gransden 2004
Last modifiedJanuary 15, 2020
Genome assembly and annotationi <p>Identifier for the genome assembly (<a href="">more...</a>)</p> GCA_000002425.2 from ENA/EMBL full

The moss Physcomitrella patens is a model organism for studying the evolution of land plants. As a Bryophyte this organism sits midway along the evolutionary path between unicellular aquatic algae and flowering land plants.

Physcomitrella patens has a very efficient mechanism of homologous recombination which can be used to create targeted genetic modifications. It also shows alternation of generations, with the haploid phase of the life-cycle being dominant. These characteristics make it an excellent experimental material. It is used to examine the key genetic changes involved in adaptation to water stress and temperature variation, associated with life on land, and the loss of flagellae, associated with aquatic life.

The reference proteome for Physcomitrella patens is derived from the genome published in 2008 ( ). The genome has a haploid chromosome number of 27 containing nearly 500 Mb, with 35,000 protein-coding genes. Further details of the genome assembly are available at the 'Physcomitrella patens resource' (

Componentsi <p>Genomic components encoding the proteome</p>

Component nameGenome Accession(s)
Component representationProteins
Chromosome 11939
Chromosome 21773
Chromosome 31738
Chromosome 41439
Chromosome 51249
Chromosome 61346
Chromosome 71250
Chromosome 81090
Chromosome 91143
Chromosome 101157
Chromosome 111240
Chromosome 121120
Chromosome 131006
Chromosome 141260
Chromosome 151130
Chromosome 161202
Chromosome 171093
Chromosome 18894
Chromosome 19966
Chromosome 201042
Chromosome 21951
Chromosome 22950
Chromosome 23962
Chromosome 24908
Chromosome 25634
Chromosome 26671
Chromosome 27400
Unassembled WGS sequence178


  1. "The mitochondrial genome of the moss Physcomitrella patens sheds new light on mitochondrial evolution in land plants."
    Terasawa K., Odahara M., Kabeya Y., Kikugawa T., Sekine Y., Fujiwara M., Sato N.
    Mol. Biol. Evol. 24:699-709(2007) [PubMed] [Europe PMC] [Abstract]
  2. "The Physcomitrella genome reveals evolutionary insights into the conquest of land by plants."
    Rensing S.A., Lang D., Zimmer A.D., Terry A., Salamov A., Shapiro H., Nishiyama T., Perroud P.F., Lindquist E.A., Kamisugi Y., Tanahashi T., Sakakibara K., Fujita T., Oishi K., Shin-I T., Kuroki Y., Toyoda A., Suzuki Y.
    Boore J.L.
    Science 319:64-69(2008) [PubMed] [Europe PMC] [Abstract]
  3. "The Physcomitrella patens chromosome-scale assembly reveals moss genome structure and evolution."
    Lang D., Ullrich K.K., Murat F., Fuchs J., Jenkins J., Haas F.B., Piednoel M., Gundlach H., Van Bel M., Meyberg R., Vives C., Morata J., Symeonidi A., Hiss M., Muchero W., Kamisugi Y., Saleh O., Blanc G.
    Rensing S.A.
    Plant Journal 93:515-533(2018) [PubMed] [Europe PMC] [Abstract]
  4. "Complete chloroplast DNA sequence of the moss Physcomitrella patens: evidence for the loss and relocation of rpoA from the chloroplast to the nucleus."
    Sugiura C., Kobayashi Y., Setsuyuki A., Sugita C., Sugita M.
    Nucleic Acids Res. 31:5324-5331(2003) [PubMed] [Europe PMC] [Abstract]
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