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StatusReference proteome
Gene counti <p>This is the total number of unique genes found in the proteome set, algorithmically computed. For each gene, a single representative protein sequence is chosen from the proteome. Where possible, reviewed (Swiss-Prot) protein sequences are chosen as the representatives.</p> - Download one protein sequence per gene (FASTA)
Proteome IDi <p>The proteome identifier (UPID) is the unique identifier assigned to the set of proteins that constitute the <a href="">proteome</a>. It consists of the characters ‘UP’ followed by 9 digits, is stable across releases and can therefore be used to cite a UniProt proteome.<p><a href='/help/proteome_id' target='_top'>More...</a></p>UP000006727
Straincv. Gransden 2004
Last modifiedOctober 26, 2018
Genome assembly and annotationi GCA_000002425.2 from ENA/EMBL

The moss Physcomitrella patens is a model organism for studying the evolution of land plants. As a Bryophyte this organism sits midway along the evolutionary path between unicellular aquatic algae and flowering land plants.

Physcomitrella patens has a very efficient mechanism of homologous recombination which can be used to create targeted genetic modifications. It also shows alternation of generations, with the haploid phase of the life-cycle being dominant. These characteristics make it an excellent experimental material. It is used to examine the key genetic changes involved in adaptation to water stress and temperature variation, associated with life on land, and the loss of flagellae, associated with aquatic life.

The reference proteome for Physcomitrella patens is derived from the genome published in 2008 ( ). The genome has a haploid chromosome number of 27 containing nearly 500 Mb, with 35,000 protein-coding genes. Further details of the genome assembly are available at the 'Physcomitrella patens resource' (

Componentsi <p>Genomic components encoding the proteome</p>

Component nameGenome Accession(s)
Chromosome 11947
Chromosome 21783
Chromosome 31741
Chromosome 41450
Chromosome 51255
Chromosome 61351
Chromosome 71256
Chromosome 81094
Chromosome 91145
Chromosome 101160
Chromosome 111242
Chromosome 121123
Chromosome 131008
Chromosome 141265
Chromosome 151134
Chromosome 161206
Chromosome 171094
Chromosome 18896
Chromosome 19966
Chromosome 201048
Chromosome 21953
Chromosome 22954
Chromosome 23964
Chromosome 24912
Chromosome 25635
Chromosome 26672
Chromosome 27401
Unassembled WGS sequence163


  1. "The Physcomitrella genome reveals evolutionary insights into the conquest of land by plants."
    Rensing S.A., Lang D., Zimmer A.D., Terry A., Salamov A., Shapiro H., Nishiyama T., Perroud P.F., Lindquist E.A., Kamisugi Y., Tanahashi T., Sakakibara K., Fujita T., Oishi K., Shin-I T., Kuroki Y., Toyoda A., Suzuki Y.
    Boore J.L.
    Science 2008:64-69(2008) [PubMed] [Europe PMC] [Abstract]
  2. "The Physcomitrella patens chromosome-scale assembly reveals moss genome structure and evolution."
    Lang D., Ullrich K.K., Murat F., Fuchs J., Jenkins J., Haas F.B., Piednoel M., Gundlach H., Van Bel M., Meyberg R., Vives C., Morata J., Symeonidi A., Hiss M., Muchero W., Kamisugi Y., Saleh O., Blanc G.
    Rensing S.A.
    Plant Journal 2018:515-533(2018) [PubMed] [Europe PMC] [Abstract]
  3. "Complete chloroplast DNA sequence of the moss Physcomitrella patens: evidence for the loss and relocation of rpoA from the chloroplast to the nucleus."
    Sugiura C., Kobayashi Y., Setsuyuki A., Sugita C., Sugita M.
    Nucleic Acids Res. 2003:5324-5331(2003) [PubMed] [Europe PMC] [Abstract]
  4. "The mitochondrial genome of the moss Physcomitrella patens sheds new light on mitochondrial evolution in land plants."
    Terasawa K., Odahara M., Kabeya Y., Kikugawa T., Sekine Y., Fujiwara M., Sato N.
    Mol. Biol. Evol. 2007:699-709(2007) [PubMed] [Europe PMC] [Abstract]
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