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        UniProt - Swiss-Prot Protein Knowledgebase
        SIB Swiss Institute of Bioinformatics; Geneva, Switzerland
        European Bioinformatics Institute (EBI); Hinxton, United Kingdom
        Protein Information Resource (PIR); Washington DC, USA

Description: Human chromosome 4: entries, gene names and
             cross-references to MIM
Name:        humchr04.txt
Release:     2021_04 of 29-Sep-2021


This documents lists all the human protein sequence entries whose genes
are known to be encoded on chromosome 4 in this release of UniProtKB/Swiss-Prot.

Number of UniProtKB/Swiss-Prot entries encoded on chromosome 4: 768

Reference for the chromosome sequence:
Nature 434:724-731(2005).
[Pubmed: 15815621] [Article from publisher]

For more information on chromosome 4 see:

Ensembl :
NCBI    :
OMIM    :
DOE     :

______________ _______________ ______________________ ______ ______________________
Gene           Chromosomal     Swiss-Prot             MIM    Description
name           position        AC        Entry name   code
______________ _______________ ______________________ ______ ______________________
AADAT         4q33            Q8N5Z0     AADAT_HUMAN 611754 Kynurenine/alpha-aminoadipate aminotransferase, mitochondrial precursor (KAT/AadAT) (2-aminoadipate aminotransferase) (2-aminoadipate transaminase) (EC (Alpha-aminoadipate aminotransferase) (AadAT) (Kynurenine aminotransferase II) (Kynurenine--oxoglutarate aminotransferase II) (Kynurenine--oxoglutarate transaminase 2) (EC (Kynurenine--oxoglutarate transaminase II) [KAT2] [KYAT2]
AASDH         4q12            Q4L235     ACSF4_HUMAN 614365 Beta-alanine-activating enzyme (EC 6.2.1.-) (Acyl-CoA synthetase family member 4) (Protein NRPS998) [ACSF4] [U26] [HSPC318]
ABCE1         4q31            P61221     ABCE1_HUMAN 601213 ATP-binding cassette sub-family E member 1 (2'-5'-oligoadenylate-binding protein) (HuHP68) (RNase L inhibitor) (Ribonuclease 4 inhibitor) (RNS4I) [RLI] [RNASEL1] [RNASELI] [RNS4I] [OK/SW-cl.40]
ABCG2         4q22            Q9UNQ0     ABCG2_HUMAN 603756 Broad substrate specificity ATP-binding cassette transporter ABCG2 (EC (ATP-binding cassette sub-family G member 2) (Breast cancer resistance protein) (CDw338) (Mitoxantrone resistance-associated protein) (Placenta-specific ATP-binding cassette transporter) (Urate exporter) (CD338 antigen) [ABCP] [BCRP] [BCRP1] [MXR]
ABHD18        4q28.2          Q0P651     ABD18_HUMAN        Protein ABHD18 precursor (Alpha/beta hydrolase domain-containing protein 18) (Abhydrolase domain-containing protein 18) [C4orf29]
ABLIM2        4p16.1          Q6H8Q1     ABLM2_HUMAN 612544 Actin-binding LIM protein 2 (abLIM-2) (Actin-binding LIM protein family member 2) [KIAA1808]
ABRAXAS1      4q21.23         Q6UWZ7     ABRX1_HUMAN 611143 BRCA1-A complex subunit Abraxas 1 (Coiled-coil domain-containing protein 98) (Protein FAM175A) [ABRA1] [CCDC98] [FAM175A] [UNQ496/PRO1013]
ACOT7L        4p14            Q6ZUV0     BACHL_HUMAN 611963 Putative cytosolic acyl coenzyme A thioester hydrolase-like (EC (Acyl-CoA thioesterase 7-like) [BACHL]
ACOX3         4p15.3          O15254     ACOX3_HUMAN 603402 Peroxisomal acyl-coenzyme A oxidase 3 (EC (Branched-chain acyl-CoA oxidase) (BRCACox) (Pristanoyl-CoA oxidase) [BRCOX] [PRCOX]
ACSL1         4q34-q35        P33121     ACSL1_HUMAN 152425 Long-chain-fatty-acid--CoA ligase 1 (EC (Acyl-CoA synthetase 1) (ACS1) (Arachidonate--CoA ligase) (EC (Long-chain acyl-CoA synthetase 1) (LACS 1) (Long-chain acyl-CoA synthetase 2) (LACS 2) (Long-chain fatty acid-CoA ligase 2) (Palmitoyl-CoA ligase 1) (Palmitoyl-CoA ligase 2) (Phytanate--CoA ligase) (EC [FACL1] [FACL2] [LACS] [LACS1] [LACS2]
ADAD1         4q27            Q96M93     ADAD1_HUMAN 614130 Adenosine deaminase domain-containing protein 1 (Testis nuclear RNA-binding protein) [TENR]
ADAM29        4q34.2-qter     Q9UKF5     ADA29_HUMAN 604778 Disintegrin and metalloproteinase domain-containing protein 29 precursor (ADAM 29) (Cancer/testis antigen 73) (CT73)
ADAMTS3       4q21.1-q21.22   O15072     ATS3_HUMAN  605011 A disintegrin and metalloproteinase with thrombospondin motifs 3 precursor (EC 3.4.24.-) (ADAM-TS 3) (ADAM-TS3) (ADAMTS-3) (Procollagen II N-proteinase) (PC II-NP) (Procollagen II amino propeptide-processing enzyme) [KIAA0366]
ADD1          4p16.3          P35611     ADDA_HUMAN  102680 Alpha-adducin (Erythrocyte adducin subunit alpha) [ADDA]
ADGRA3        4p15.31         Q8IWK6     AGRA3_HUMAN 612303 Adhesion G protein-coupled receptor A3 precursor (G-protein coupled receptor 125) [GPR125] [UNQ556/PRO1113]
ADGRL3        4q13.1          Q9HAR2     AGRL3_HUMAN 616417 Adhesion G protein-coupled receptor L3 precursor (Calcium-independent alpha-latrotoxin receptor 3) (CIRL-3) (Latrophilin-3) (Lectomedin-3) [KIAA0768] [LEC3] [LPHN3]
ADH1A         4q21-q23        P07327     ADH1A_HUMAN 103700 Alcohol dehydrogenase 1A (EC (Alcohol dehydrogenase subunit alpha) [ADH1]
ADH1B         4q21-q23        P00325     ADH1B_HUMAN 103720 All-trans-retinol dehydrogenase [NAD(+)] ADH1B (EC (Alcohol dehydrogenase 1B) (Alcohol dehydrogenase subunit beta) [ADH2]
ADH1C         4q21-q23        P00326     ADH1G_HUMAN 103730 Alcohol dehydrogenase 1C (EC (Alcohol dehydrogenase subunit gamma) [ADH3]
ADH4          4q22            P08319     ADH4_HUMAN  103740 All-trans-retinol dehydrogenase [NAD(+)] ADH4 (EC (Alcohol dehydrogenase 4) (Alcohol dehydrogenase class II pi chain)
ADH5          4q21-q24        P11766     ADHX_HUMAN  103710 Alcohol dehydrogenase class-3 (EC (Alcohol dehydrogenase 5) (Alcohol dehydrogenase class chi chain) (Alcohol dehydrogenase class-III) (Glutathione-dependent formaldehyde dehydrogenase) (EC 1.1.1.-) (FALDH) (FDH) (GSH-FDH) (S-(hydroxymethyl)glutathione dehydrogenase) (EC [ADHX] [FDH]
ADH6          4q23            P28332     ADH6_HUMAN  103735 Alcohol dehydrogenase 6 (EC
ADH7          4q23-q24        P40394     ADH7_HUMAN  600086 All-trans-retinol dehydrogenase [NAD(+)] ADH7 (EC (Alcohol dehydrogenase class 4 mu/sigma chain) (EC (Alcohol dehydrogenase class IV mu/sigma chain) (Gastric alcohol dehydrogenase) (Omega-hydroxydecanoate dehydrogenase ADH7) (EC (Retinol dehydrogenase)
ADRA2C        4p16.1          P18825     ADA2C_HUMAN 104250 Alpha-2C adrenergic receptor (Alpha-2 adrenergic receptor subtype C4) (Alpha-2C adrenoreceptor) (Alpha-2C adrenoceptor) (Alpha-2CAR) [ADRA2L2] [ADRA2RL2]
AFAP1         4p16            Q8N556     AFAP1_HUMAN 608252 Actin filament-associated protein 1 (110 kDa actin filament-associated protein) (AFAP-110) [AFAP]
AFF1          4q21            P51825     AFF1_HUMAN  159557 AF4/FMR2 family member 1 (ALL1-fused gene from chromosome 4 protein) (Protein AF-4) (Protein FEL) (Proto-oncogene AF4) [AF4] [FEL] [MLLT2] [PBM1]
AFM           4q11-q13        P43652     AFAM_HUMAN  104145 Afamin precursor (Alpha-albumin) (Alpha-Alb) [ALB2] [ALBA]
AFP           4q11-q13        P02771     FETA_HUMAN  104150 Alpha-fetoprotein precursor (Alpha-1-fetoprotein) (Alpha-fetoglobulin) [HPAFP]
AGA           4q32-q33        P20933     ASPG_HUMAN  613228 N(4)-(beta-N-acetylglucosaminyl)-L-asparaginase precursor (EC (Aspartylglucosaminidase) (Glycosylasparaginase) (N4-(N-acetyl-beta-glucosaminyl)-L-asparagine amidase) [Contains: Glycosylasparaginase alpha chain; Glycosylasparaginase beta chain]
AIMP1         4q24            Q12904     AIMP1_HUMAN 603605 Aminoacyl tRNA synthase complex-interacting multifunctional protein 1 (Multisynthase complex auxiliary component p43) [Contains: Endothelial monocyte-activating polypeptide 2 (EMAP-2) (Endothelial monocyte-activating polypeptide II) (EMAP-II) (Small inducible cytokine subfamily E member 1)] [EMAP2] [SCYE1]
ALB           4q11-q13        P02768     ALBU_HUMAN  103600 Albumin precursor [GIG20] [GIG42] [PRO0903] [PRO1708] [PRO2044] [PRO2619] [PRO2675] [UNQ696/PRO1341]
ALPK1         4q25            Q96QP1     ALPK1_HUMAN 607347 Alpha-protein kinase 1 (EC (Chromosome 4 kinase) (Lymphocyte alpha-protein kinase) [KIAA1527] [LAK]
AMBN          4q21            Q9NP70     AMBN_HUMAN  601259 Ameloblastin precursor
AMTN          4q13.3          Q6UX39     AMTN_HUMAN  610912 Amelotin precursor [UNQ689/PRO1329]
ANAPC10       4q31            Q9UM13     APC10_HUMAN 613745 Anaphase-promoting complex subunit 10 (APC10) (Cyclosome subunit 10) [APC10]
ANAPC4        4p15.2          Q9UJX5     APC4_HUMAN  606947 Anaphase-promoting complex subunit 4 (APC4) (Cyclosome subunit 4) [APC4]
ANK2          4q25-q27        Q01484     ANK2_HUMAN  106410 Ankyrin-2 (ANK-2) (Ankyrin-B) (Brain ankyrin) (Non-erythroid ankyrin) [ANKB]
ANKRD17       4q13.3          O75179     ANR17_HUMAN 615929 Ankyrin repeat domain-containing protein 17 (Gene trap ankyrin repeat protein) (Serologically defined breast cancer antigen NY-BR-16) [GTAR] [KIAA0697]
ANKRD37       4q35.1          Q7Z713     ANR37_HUMAN 619021 Ankyrin repeat domain-containing protein 37 (Low-density lipoprotein receptor-related protein 2-binding protein) (hLrp2bp) [LPR2BP]
ANKRD50       4q26            Q9ULJ7     ANR50_HUMAN        Ankyrin repeat domain-containing protein 50 [KIAA1223]
ANP32C        4q32.3          O43423     AN32C_HUMAN 606877 Acidic leucine-rich nuclear phosphoprotein 32 family member C (Phosphoprotein 32-related protein 1) (Tumorigenic protein pp32r1) [PP32R1]
ANTXR2        4q21.3          P58335     ANTR2_HUMAN 608041 Anthrax toxin receptor 2 precursor (Capillary morphogenesis gene 2 protein) (CMG-2) [CMG2]
ANXA10        4q33            Q9UJ72     ANX10_HUMAN 608008 Annexin A10 (Annexin-10) (Annexin-14) [ANX14]
ANXA3         4q13-q22        P12429     ANXA3_HUMAN 106490 Annexin A3 (35-alpha calcimedin) (Annexin III) (Annexin-3) (Inositol 1,2-cyclic phosphate 2-phosphohydrolase) (Lipocortin III) (Placental anticoagulant protein III) (PAP-III) [ANX3]
ANXA5         4q28-q32        P08758     ANXA5_HUMAN 131230 Annexin A5 (Anchorin CII) (Annexin V) (Annexin-5) (Calphobindin I) (CBP-I) (Endonexin II) (Lipocortin V) (Placental anticoagulant protein 4) (PP4) (Placental anticoagulant protein I) (PAP-I) (Thromboplastin inhibitor) (Vascular anticoagulant-alpha) (VAC-alpha) [ANX5] [ENX2] [PP4]
AP1AR         4q25            Q63HQ0     AP1AR_HUMAN 610851 AP-1 complex-associated regulatory protein (2c18) (Adaptor-related protein complex 1-associated regulatory protein) (Gamma-1-adaptin brefeldin A resistance protein) (GBAR) (Gamma-BAR) (Gamma-A1-adaptin and kinesin interactor) (Gadkin) [C4orf16] [PRO0971]
APBB2         4p13            Q92870     APBB2_HUMAN 602710 Amyloid beta precursor protein binding family B member 2 (Amyloid-beta (A4) precursor protein-binding family B member 2) (Protein Fe65-like 1) [FE65L] [FE65L1]
APELA         4q32            P0DMC3     ELA_HUMAN   615594 Apelin receptor early endogenous ligand precursor (Protein Elabela) (ELA) (Protein Toddler) [ELA] [TDL]
ARAP2         4p14            Q8WZ64     ARAP2_HUMAN 606645 Arf-GAP with Rho-GAP domain, ANK repeat and PH domain-containing protein 2 (Centaurin-delta-1) (Cnt-d1) (Protein PARX) [CENTD1] [KIAA0580]
AREG          4q13.3          P15514     AREG_HUMAN  104640 Amphiregulin precursor (AR) (Colorectum cell-derived growth factor) (CRDGF) [AREGB] [SDGF]
ARFIP1        4q31.23         P53367     ARFP1_HUMAN 605928 Arfaptin-1 (ADP-ribosylation factor-interacting protein 1)
ARHGAP10      4q31.23         A1A4S6     RHG10_HUMAN 609746 Rho GTPase-activating protein 10 (GTPase regulator associated with focal adhesion kinase 2) (Graf-related protein 2) (Rho-type GTPase-activating protein 10) [GRAF2]
ARHGAP24      4q21.23-q21.3   Q8N264     RHG24_HUMAN 610586 Rho GTPase-activating protein 24 (Filamin-A-associated RhoGAP) (FilGAP) (RAC1- and CDC42-specific GTPase-activating protein of 72 kDa) (RC-GAP72) (Rho-type GTPase-activating protein 24) (RhoGAP of 73 kDa) (Sarcoma antigen NY-SAR-88) (p73RhoGAP) [FILGAP]
ARHGEF38      4q24            Q9NXL2     ARH38_HUMAN        Rho guanine nucleotide exchange factor 38
ARL9          4q12            Q6T311     ARL9_HUMAN  612405 ADP-ribosylation factor-like protein 9
ARSJ          4q26            Q5FYB0     ARSJ_HUMAN  610010 Arylsulfatase J precursor (EC 3.1.6.-) (ASJ) [UNQ372/PRO708]
ART3          4p14-p15.1      Q13508     NAR3_HUMAN  603086 Ecto-ADP-ribosyltransferase 3 precursor (EC (ADP-ribosyltransferase C2 and C3 toxin-like 3) (ARTC3) (Mono(ADP-ribosyl)transferase 3) (NAD(P)(+)--arginine ADP-ribosyltransferase 3) [TMART]
ASB5          4q34.1          Q8WWX0     ASB5_HUMAN  615050 Ankyrin repeat and SOCS box protein 5 (ASB-5)
ASIC5         4q32.1          Q9NY37     ASIC5_HUMAN 616693 Acid-sensing ion channel 5 (ASIC5) (Amiloride-sensitive cation channel 5) (Human intestine Na(+) channel) (HINaC) [ACCN5] [HINAC]
ATOH1         4q22            Q92858     ATOH1_HUMAN 601461 Protein atonal homolog 1 (Class A basic helix-loop-helix protein 14) (bHLHa14) (Helix-loop-helix protein hATH-1) (hATH1) [ATH1] [BHLHA14]
ATP10D        4p11            Q9P241     AT10D_HUMAN        Phospholipid-transporting ATPase VD (EC (ATPase class V type 10D) (P4-ATPase flippase complex alpha subunit ATP10D) [ATPVD] [KIAA1487]
ATP5ME        4p              P56385     ATP5I_HUMAN 601519 ATP synthase subunit e, mitochondrial (ATPase subunit e) (ATP synthase membrane subunit e) [Contains: ATP synthase subunit e, mitochondrial, N-terminally processed] [ATP5I] [ATP5K]
ATP8A1        4p12-p14        Q9Y2Q0     AT8A1_HUMAN 609542 Phospholipid-transporting ATPase IA (EC (ATPase class I type 8A member 1) (Chromaffin granule ATPase II) (P4-ATPase flippase complex alpha subunit ATP8A1) [ATPIA]
BANK1         4q24            Q8NDB2     BANK1_HUMAN 610292 B-cell scaffold protein with ankyrin repeats
BBS12         4q27            Q6ZW61     BBS12_HUMAN 610683 Bardet-Biedl syndrome 12 protein [C4orf24]
BBS7          4q27            Q8IWZ6     BBS7_HUMAN  607590 Bardet-Biedl syndrome 7 protein (BBS2-like protein 1) [BBS2L1]
BDH2          4q24            Q9BUT1     BDH2_HUMAN         3-hydroxybutyrate dehydrogenase type 2 (EC 1.1.1.-) (EC (Dehydrogenase/reductase SDR family member 6) (Oxidoreductase UCPA) (R-beta-hydroxybutyrate dehydrogenase) (Short chain dehydrogenase/reductase family 15C member 1) [DHRS6] [SDR15C1] [UNQ6308/PRO20933]
BEND4         4p13            Q6ZU67     BEND4_HUMAN        BEN domain-containing protein 4 (Coiled-coil domain-containing protein 4) [CCDC4]
BLOC1S4       4p16.2          Q9NUP1     BL1S4_HUMAN 605695 Biogenesis of lysosome-related organelles complex 1 subunit 4 (BLOC-1 subunit 4) (Protein cappuccino homolog) [CNO]
BMP2K         4q21.23         Q9NSY1     BMP2K_HUMAN 617648 BMP-2-inducible protein kinase (EC (BIKe) [BIKE] [HRIHFB2017]
BMP3          4p14-q21        P12645     BMP3_HUMAN  112263 Bone morphogenetic protein 3 precursor (BMP-3) (Bone morphogenetic protein 3A) (BMP-3A) (Osteogenin) [BMP3A]
BMPR1B        4q23-q24        O00238     BMR1B_HUMAN 603248 Bone morphogenetic protein receptor type-1B precursor (EC (BMP type-1B receptor) (BMPR-1B) (CDw293 antigen)
BOD1L1        4p16.1          Q8NFC6     BD1L1_HUMAN 616746 Biorientation of chromosomes in cell division protein 1-like 1 [BOD1L] [FAM44A] [KIAA1327]
BST1          4p15            Q10588     BST1_HUMAN  600387 ADP-ribosyl cyclase/cyclic ADP-ribose hydrolase 2 precursor (EC (ADP-ribosyl cyclase 2) (Bone marrow stromal antigen 1) (BST-1) (Cyclic ADP-ribose hydrolase 2) (cADPr hydrolase 2) (CD157 antigen)
BTC           4q13-q21        P35070     BTC_HUMAN   600345 Probetacellulin precursor [Contains: Betacellulin (BTC)]
C1QTNF7       4p15            Q9BXJ2     C1QT7_HUMAN        Complement C1q tumor necrosis factor-related protein 7 precursor [CTRP7]
C4orf17       4q23            Q53FE4     CD017_HUMAN        Uncharacterized protein C4orf17
C4orf19       4p14            Q8IY42     CD019_HUMAN        Uncharacterized protein C4orf19
C4orf3        4q26            Q8WVX3     CD003_HUMAN        Uncharacterized protein C4orf3 (Hepatitis C virus F protein-transactivated protein 1) (HCV F-transactivated protein 1)
C4orf33       4q28.2          Q8N1A6     CD033_HUMAN        UPF0462 protein C4orf33
C4orf36       4q21.3          Q96KX1     CD036_HUMAN        Uncharacterized protein C4orf36
C4orf45       4q32.1          Q96LM5     CD045_HUMAN        Uncharacterized protein C4orf45
C4orf46       4q32.1          Q504U0     CD046_HUMAN 616210 Renal cancer differentiation gene 1 protein [RCDG1]
C4orf47       4q35.1          A7E2U8     CD047_HUMAN        UPF0602 protein C4orf47 [Chr4_1746]
C4orf48       4p16.3          Q5BLP8     CD048_HUMAN 614690 Neuropeptide-like protein C4orf48 precursor [Chr4_55]
C4orf50       4p16.1          Q6ZRC1     CD050_HUMAN        Uncharacterized protein C4orf50
C4orf51       4q31.21         C9J302     CD051_HUMAN        Uncharacterized protein C4orf51
C4orf54       4q23            D6RIA3     CD054_HUMAN 617881 Uncharacterized protein C4orf54 (Familial obliterative portal venopathy) [FOPV]
CABS1         4q13.3          Q96KC9     CABS1_HUMAN 618600 Calcium-binding and spermatid-specific protein 1 (Testis development protein NYD-SP26) [C4orf35]
CAMK2D        4q25            Q13557     KCC2D_HUMAN 607708 Calcium/calmodulin-dependent protein kinase type II subunit delta (EC (CaM kinase II subunit delta) (CaMK-II subunit delta) [CAMKD]
CASP3         4q33-q35.1      P42574     CASP3_HUMAN 600636 Caspase-3 precursor (EC (CASP-3) (Apopain) (Cysteine protease CPP32) (CPP-32) (Protein Yama) (SREBP cleavage activity 1) (SCA-1) [Contains: Caspase-3 subunit p17; Caspase-3 subunit p12] [CPP32]
CASP6         4q25            P55212     CASP6_HUMAN 601532 Caspase-6 precursor (EC (CASP-6) (Apoptotic protease Mch-2) [Contains: Caspase-6 subunit p18; Caspase-6 subunit p11] [MCH2]
CBR4          4q32.3          Q8N4T8     CBR4_HUMAN         3-oxoacyl-[acyl-carrier-protein] reductase (EC (3-ketoacyl-[acyl-carrier-protein] reductase beta subunit) (KAR beta subunit) (Carbonyl reductase family member 4) (CBR4) (Quinone reductase CBR4) (EC (Short chain dehydrogenase/reductase family 45C member 1) [SDR45C1]
CC2D2A        4p15.33         Q9P2K1     C2D2A_HUMAN 612013 Coiled-coil and C2 domain-containing protein 2A [KIAA1345]
CCDC110       4q35.1          Q8TBZ0     CC110_HUMAN 609488 Coiled-coil domain-containing protein 110 (Cancer/testis antigen 52) (CT52) (Cancer/testis antigen KM-HN-1) [KMHN1]
CCDC149       4p15.2          Q6ZUS6     CC149_HUMAN        Coiled-coil domain-containing protein 149
CCDC158       4q21.1          Q5M9N0     CD158_HUMAN        Coiled-coil domain-containing protein 158
CCDC96        4p16.1          Q2M329     CCD96_HUMAN 619347 Coiled-coil domain-containing protein 96
CCKAR         4p15.1-p15.2    P32238     CCKAR_HUMAN 118444 Cholecystokinin receptor type A (CCK-A receptor) (CCK-AR) (Cholecystokinin-1 receptor) (CCK1-R) [CCKRA]
CCNA2         4q27            P20248     CCNA2_HUMAN 123835 Cyclin-A2 (Cyclin-A) (Cyclin A) [CCN1] [CCNA]
CCNG2         4q13.3          Q16589     CCNG2_HUMAN 603203 Cyclin-G2
CCNI          4q13.3          Q14094     CCNI_HUMAN  618783 Cyclin-I
CCSER1        4q22.1          Q9C0I3     CCSE1_HUMAN 618934 Serine-rich coiled-coil domain-containing protein 1 (Coiled-coil serine-rich protein 1) [FAM190A] [KIAA1680]
CD38          4p15            P28907     CD38_HUMAN  107270 ADP-ribosyl cyclase/cyclic ADP-ribose hydrolase 1 (EC (2'-phospho-ADP-ribosyl cyclase) (2'-phospho-ADP-ribosyl cyclase/2'-phospho-cyclic-ADP-ribose transferase) (EC (2'-phospho-cyclic-ADP-ribose transferase) (ADP-ribosyl cyclase 1) (ADPRC 1) (Cyclic ADP-ribose hydrolase 1) (cADPr hydrolase 1) (T10) (CD38 antigen)
CDKL2         4q21            Q92772     CDKL2_HUMAN 603442 Cyclin-dependent kinase-like 2 (EC (Protein kinase p56 KKIAMRE) (Serine/threonine-protein kinase KKIAMRE)
CDKN2AIP      4q35.1          Q9NXV6     CARF_HUMAN  615914 CDKN2A-interacting protein (Collaborator of ARF) [CARF]
CDS1          4q21            Q92903     CDS1_HUMAN  603548 Phosphatidate cytidylyltransferase 1 (EC (CDP-DAG synthase 1) (CDP-DG synthase 1) (CDP-diacylglycerol synthase 1) (CDS 1) (CDP-diglyceride pyrophosphorylase 1) (CDP-diglyceride synthase 1) (CTP:phosphatidate cytidylyltransferase 1) [CDS]
CENPC         4q12-q13.3      Q03188     CENPC_HUMAN 117141 Centromere protein C (CENP-C) (Centromere autoantigen C) (Centromere protein C 1) (CENP-C 1) (Interphase centromere complex protein 7) [CENPC1] [ICEN7]
CENPE         4q24-q25        Q02224     CENPE_HUMAN 117143 Centromere-associated protein E precursor (Centromere protein E) (CENP-E) (Kinesin-7) (Kinesin-related protein CENPE)
CENPU         4q35.1          Q71F23     CENPU_HUMAN 611511 Centromere protein U (CENP-U) (Centromere protein of 50 kDa) (CENP-50) (Interphase centromere complex protein 24) (KSHV latent nuclear antigen-interacting protein 1) (MLF1-interacting protein) (Polo-box-interacting protein 1) [ICEN24] [KLIP1] [MLF1IP] [PBIP1]
CEP135        4q12-q13.1      Q66GS9     CP135_HUMAN 611423 Centrosomal protein of 135 kDa (Cep135) (Centrosomal protein 4) [CEP4] [KIAA0635]
CEP170P1      4q26            Q96L14     C170L_HUMAN        Cep170-like protein (CEP170 pseudogene 1) [CEP170L] [KIAA0470L]
CEP44         4q34            Q9C0F1     CEP44_HUMAN        Centrosomal protein of 44 kDa (Cep44) (HBV PreS1-transactivated protein 3) (PS1TP3) [KIAA1712]
CFAP299       4q21.21         Q6V702     CF299_HUMAN        Cilia- and flagella-associated protein 299 [C4orf22]
CFAP97        4q35.1          Q9P2B7     CFA97_HUMAN 616047 Cilia- and flagella-associated protein 97 [KIAA1430]
CFAP99        4p16.3          D6REC4     CFA99_HUMAN        Cilia- and flagella-associated protein 99
CFI           4q25            P05156     CFAI_HUMAN  217030 Complement factor I precursor (EC (C3B/C4B inactivator) [Contains: Complement factor I heavy chain; Complement factor I light chain] [IF]
CHIC2         4q12            Q9UKJ5     CHIC2_HUMAN 604332 Cysteine-rich hydrophobic domain-containing protein 2 (BrX-like translocated in leukemia) [BTL]
CHRNA9        4p14            Q9UGM1     ACHA9_HUMAN 605116 Neuronal acetylcholine receptor subunit alpha-9 precursor (Nicotinic acetylcholine receptor subunit alpha-9) (NACHR alpha-9) [NACHRA9]
CISD2         4q24            Q8N5K1     CISD2_HUMAN 611507 CDGSH iron-sulfur domain-containing protein 2 (Endoplasmic reticulum intermembrane small protein) (MitoNEET-related 1 protein) (Miner1) (Nutrient-deprivation autophagy factor-1) (NAF-1) [CDGSH2] [ERIS] [ZCD2]
CLCN3         4q33            P51790     CLCN3_HUMAN 600580 H(+)/Cl(-) exchange transporter 3 (Chloride channel protein 3) (ClC-3) (Chloride transporter ClC-3)
CLDN22        4q35.1          Q8N7P3     CLD22_HUMAN        Claudin-22
CLDN24        4q35.1          A6NM45     CLD24_HUMAN        Putative claudin-24 (Claudin-21) [CLDN21]
CLGN          4q28.3-q31.1    O14967     CLGN_HUMAN  601858 Calmegin precursor
CLNK          4p16.1          Q7Z7G1     CLNK_HUMAN  611434 Cytokine-dependent hematopoietic cell linker (Mast cell immunoreceptor signal transducer) [MIST]
CLOCK         4q12            O15516     CLOCK_HUMAN 601851 Circadian locomoter output cycles protein kaput (EC (hCLOCK) (Class E basic helix-loop-helix protein 8) (bHLHe8) [BHLHE8] [KIAA0334]
CLRN2         4p15.33         A0PK11     CLRN2_HUMAN 618988 Clarin-2
CNGA1         4p13-q14        P29973     CNGA1_HUMAN 123825 cGMP-gated cation channel alpha-1 (Cyclic nucleotide-gated cation channel 1) (Cyclic nucleotide-gated channel alpha-1) (CNG channel alpha-1) (CNG-1) (CNG1) (Cyclic nucleotide-gated channel, photoreceptor) (Rod photoreceptor cGMP-gated channel subunit alpha) [CNCG] [CNCG1]
CNOT6L        4q21.1          Q96LI5     CNO6L_HUMAN 618069 CCR4-NOT transcription complex subunit 6-like (EC (Carbon catabolite repressor protein 4 homolog B) [CCR4B]
COL25A1       4q25            Q9BXS0     COPA1_HUMAN 610004 Collagen alpha-1(XXV) chain (Alzheimer disease amyloid-associated protein) (AMY) (CLAC-P) [Contains: Collagen-like Alzheimer amyloid plaque component (CLAC)]
COL25A1-DT    4q25            Q6ZST2     COLDT_HUMAN        Putative uncharacterized protein COL25A1-DT (COL25A1 divergent transcript) (Zinc finger CCHC domain-containing protein 23) [ZCCHC23]
COMMD8        4p12            Q9NX08     COMD8_HUMAN 616656 COMM domain-containing protein 8 [MDS022]
COPS4         4q21.3          Q9BT78     CSN4_HUMAN  616008 COP9 signalosome complex subunit 4 (SGN4) (Signalosome subunit 4) (JAB1-containing signalosome subunit 4) [CSN4]
COQ2          4q21.23         Q96H96     COQ2_HUMAN  609825 4-hydroxybenzoate polyprenyltransferase, mitochondrial precursor (EC (4-HB polyprenyltransferase) (4-hydroxybenzoate decaprenyltransferase) (COQ2 homolog) (hCOQ2) (Para-hydroxybenzoate--polyprenyltransferase) (PHB:PPT) (PHB:polyprenyltransferase) [CL640]
CORIN         4p12-p13        Q9Y5Q5     CORIN_HUMAN 605236 Atrial natriuretic peptide-converting enzyme (EC 3.4.21.-) (Corin) (Heart-specific serine proteinase ATC2) (Pro-ANP-converting enzyme) (Transmembrane protease serine 10) [Contains: Atrial natriuretic peptide-converting enzyme, N-terminal propeptide; Atrial natriuretic peptide-converting enzyme, activated protease fragment; Atrial natriuretic peptide-converting enzyme, 180 kDa soluble fragment; Atrial natriuretic peptide-converting enzyme, 160 kDa soluble fragment; Atrial natriuretic peptide-converting enzyme, 100 kDa soluble fragment] [CRN] [TMPRSS10]
COX18         4q13.3          Q8N8Q8     COX18_HUMAN 610428 Cytochrome c oxidase assembly protein COX18, mitochondrial precursor (COX18Hs) (Cytochrome c oxidase assembly protein 18) [OXA1L2]
COX7A2P2      4q22.3          O60397     COX7S_HUMAN 123997 Putative cytochrome c oxidase subunit 7A3, mitochondrial precursor (Cytochrome c oxidase subunit VIIa 3) [COX7A3] [COX7AL2] [COX7AP2]
COX7B2        4p12            Q8TF08     CX7B2_HUMAN 609811 Cytochrome c oxidase subunit 7B2, mitochondrial precursor (Cytochrome c oxidase polypeptide VIIb2)
CPE           4q32.3          P16870     CBPE_HUMAN  114855 Carboxypeptidase E precursor (EC (CPE) (Carboxypeptidase H) (CPH) (Enkephalin convertase) (Prohormone-processing carboxypeptidase)
CPEB2         4p15.33         Q7Z5Q1     CPEB2_HUMAN 610605 Cytoplasmic polyadenylation element-binding protein 2 (CPE-BP2) (CPE-binding protein 2) (hCPEB-2)
CPLX1         4p16.3          O14810     CPLX1_HUMAN 605032 Complexin-1 (Complexin I) (CPX I) (Synaphin-2)
CPZ           4p16.1          Q66K79     CBPZ_HUMAN  603105 Carboxypeptidase Z precursor (EC 3.4.17.-) (CPZ)
CRACD         4q12            Q6ZU35     CRACD_HUMAN 618327 Capping protein-inhibiting regulator of actin dynamics (Cancer-related regulator of actin dynamics) [CRAD] [KIAA1211]
CRIPAK        4p16.3          Q8N1N5     CRPAK_HUMAN 610203 Putative protein CRIPAK (Cysteine-rich PAK1 inhibitor) (CRIPak)
CRMP1         4p15-p16.1      Q14194     DPYL1_HUMAN 602462 Dihydropyrimidinase-related protein 1 (DRP-1) (Collapsin response mediator protein 1) (CRMP-1) (Inactive dihydropyrimidinase) (Unc-33-like phosphoprotein 3) (ULIP-3) [DPYSL1] [ULIP3]
CSN1S1        4q21.1          P47710     CASA1_HUMAN 115450 Alpha-S1-casein precursor [Contains: Casoxin-D] [CASA] [CSN1]
CSN2          4pter-q21       P05814     CASB_HUMAN  115460 Beta-casein precursor [CASB]
CSN3          4q21.1          P07498     CASK_HUMAN  601695 Kappa-casein precursor [CASK] [CSN10] [CSNK]
CTBP1         4p16            Q13363     CTBP1_HUMAN 602618 C-terminal-binding protein 1 (EC 1.1.1.-) (CtBP1) [CTBP]
CTSO          4q31-q32        P43234     CATO_HUMAN  600550 Cathepsin O precursor (EC [CTSO1]
CWH43         4p12-p11        Q9H720     PG2IP_HUMAN 618561 PGAP2-interacting protein (Cell wall biogenesis protein 43 C-terminal homolog) [PGAP2IP]
CXCL1         4q21            P09341     GROA_HUMAN  155730 Growth-regulated alpha protein precursor (C-X-C motif chemokine 1) (GRO-alpha(1-73)) (Melanoma growth stimulatory activity) (MGSA) (Neutrophil-activating protein 3) (NAP-3) [Contains: GRO-alpha(4-73); GRO-alpha(5-73); GRO-alpha(6-73)] [GRO] [GRO1] [GROA] [MGSA] [SCYB1]
CXCL10        4q21            P02778     CXL10_HUMAN 147310 C-X-C motif chemokine 10 precursor (10 kDa interferon gamma-induced protein) (Gamma-IP10) (IP-10) (Small-inducible cytokine B10) [Contains: CXCL10(1-73)] [INP10] [SCYB10]
CXCL11        4q21.2          O14625     CXL11_HUMAN 604852 C-X-C motif chemokine 11 precursor (Beta-R1) (H174) (Interferon gamma-inducible protein 9) (IP-9) (Interferon-inducible T-cell alpha chemoattractant) (I-TAC) (Small-inducible cytokine B11) [ITAC] [SCYB11] [SCYB9B]
CXCL13        4q21            O43927     CXL13_HUMAN 605149 C-X-C motif chemokine 13 precursor (Angie) (B cell-attracting chemokine 1) (BCA-1) (B lymphocyte chemoattractant) (CXC chemokine BLC) (Small-inducible cytokine B13) [BCA1] [BLC] [SCYB13]
CXCL2         4q21            P19875     CXCL2_HUMAN 139110 C-X-C motif chemokine 2 precursor (Growth-regulated protein beta) (Gro-beta) (Macrophage inflammatory protein 2-alpha) (MIP2-alpha) [Contains: GRO-beta(5-73) (GRO-beta-T) (Hematopoietic synergistic factor) (HSF) (SB-251353)] [GRO2] [GROB] [MIP2A] [SCYB2]
CXCL3         4q21            P19876     CXCL3_HUMAN 139111 C-X-C motif chemokine 3 precursor (GRO-gamma(1-73)) (Growth-regulated protein gamma) (GRO-gamma) (Macrophage inflammatory protein 2-beta) (MIP2-beta) [Contains: GRO-gamma(5-73)] [GRO3] [GROG] [SCYB3]
CXCL5         4q12-q13        P42830     CXCL5_HUMAN 600324 C-X-C motif chemokine 5 precursor (ENA-78(1-78)) (Epithelial-derived neutrophil-activating protein 78) (Neutrophil-activating peptide ENA-78) (Small-inducible cytokine B5) [Contains: ENA-78(8-78); ENA-78(9-78)] [ENA78] [SCYB5]
CXCL6         4q21            P80162     CXCL6_HUMAN 138965 C-X-C motif chemokine 6 precursor (Chemokine alpha 3) (CKA-3) (Granulocyte chemotactic protein 2) (GCP-2) (Small-inducible cytokine B6) [Contains: Small-inducible cytokine B6, N-processed variant 1; Small-inducible cytokine B6, N-processed variant 2; Small-inducible cytokine B6, N-processed variant 3] [GCP2] [SCYB6]
CXCL8         4q13-q21        P10145     IL8_HUMAN   146930 Interleukin-8 precursor (IL-8) (C-X-C motif chemokine 8) (Chemokine (C-X-C motif) ligand 8) (Emoctakin) (Granulocyte chemotactic protein 1) (GCP-1) (Monocyte-derived neutrophil chemotactic factor) (MDNCF) (Monocyte-derived neutrophil-activating peptide) (MONAP) (Neutrophil-activating protein 1) (NAP-1) (Protein 3-10C) (T-cell chemotactic factor) [Contains: MDNCF-a (GCP/IL-8 protein IV) (IL8/NAP1 form I); Interleukin-8 ((Ala-IL-8)77) (GCP/IL-8 protein II) (IL-8(1-77)) (IL8/NAP1 form II) (MDNCF-b); IL-8(5-77); IL-8(6-77) ((Ser-IL-8)72) (GCP/IL-8 protein I) (IL8/NAP1 form III) (Lymphocyte-derived neutrophil-activating factor) (LYNAP) (MDNCF-c) (Neutrophil-activating factor) (NAF); IL-8(7-77) (GCP/IL-8 protein V) (IL8/NAP1 form IV); IL-8(8-77) (GCP/IL-8 protein VI) (IL8/NAP1 form V); IL-8(9-77) (GCP/IL-8 protein III) (IL8/NAP1 form VI)] [IL8]
CXCL9         4q21            Q07325     CXCL9_HUMAN 601704 C-X-C motif chemokine 9 precursor (Gamma-interferon-induced monokine) (Monokine induced by interferon-gamma) (HuMIG) (MIG) (Small-inducible cytokine B9) [CMK] [MIG] [SCYB9]
CXXC4         4q22-q24        Q9H2H0     CXXC4_HUMAN 611645 CXXC-type zinc finger protein 4 (Inhibition of the Dvl and axin complex protein) [IDAX]
CYP2U1        4q25            Q7Z449     CP2U1_HUMAN 610670 Cytochrome P450 2U1 (Long-chain fatty acid omega-monooxygenase) (EC
CYP4V2        4q35.2          Q6ZWL3     CP4V2_HUMAN 608614 Cytochrome P450 4V2 (Docosahexaenoic acid omega-hydroxylase CYP4V2) (EC (Long-chain fatty acid omega-monooxygenase) (EC
CYTL1         4p15-p16        Q9NRR1     CYTL1_HUMAN 607930 Cytokine-like protein 1 precursor (Protein C17) [C4orf4] [UNQ1942/PRO4425]
DANCR         4q12            P0C864     DANCR_HUMAN 614625 Putative uncharacterized protein DANCR (Anti-differentiation ncRNA protein) (Differentiation antagonizing non-protein coding RNA) (Small nucleolar RNA host gene protein 13) [ANCR] [KIAA0114] [SNHG13]
DAPP1         4q25-q27        Q9UN19     DAPP1_HUMAN 605768 Dual adapter for phosphotyrosine and 3-phosphotyrosine and 3-phosphoinositide (hDAPP1) (B lymphocyte adapter protein Bam32) (B-cell adapter molecule of 32 kDa) [BAM32] [HSPC066]
DCAF16        4p15.32         Q9NXF7     DCA16_HUMAN        DDB1- and CUL4-associated factor 16 [C4orf30]
DCAF4L1       4p13            Q3SXM0     DC4L1_HUMAN        DDB1- and CUL4-associated factor 4-like protein 1 (WD repeat-containing protein 21B) [WDR21B]
DCHS2         4q32.1          Q6V1P9     PCD23_HUMAN 612486 Protocadherin-23 (Cadherin-27) (Cadherin-like protein CDHJ) (Cadherin-like protein VR8) (Protein dachsous homolog 2) (Protocadherin PCDHJ) [CDH27] [CDHJ] [PCDH23] [PCDHJ]
DCK           4q13.3-q21.1    P27707     DCK_HUMAN   125450 Deoxycytidine kinase (EC (dCK) (Deoxyadenosine kinase) (EC (Deoxyguanosine kinase) (EC
DCLK2         4q31.3          Q8N568     DCLK2_HUMAN 613166 Serine/threonine-protein kinase DCLK2 (EC (CaMK-like CREB regulatory kinase 2) (CL2) (CLICK-II) (CLICK2) (Doublecortin domain-containing protein 3B) (Doublecortin-like and CAM kinase-like 2) (Doublecortin-like kinase 2) [DCAMKL2] [DCDC3B] [DCK2]
DCTD          4q35.1          P32321     DCTD_HUMAN  607638 Deoxycytidylate deaminase (EC (dCMP deaminase)
DCUN1D4       4q11            Q92564     DCNL4_HUMAN 612977 DCN1-like protein 4 (DCNL4) (DCUN1 domain-containing protein 4) (Defective in cullin neddylation protein 1-like protein 4) [KIAA0276]
DDIT4L        4q23            Q96D03     DDT4L_HUMAN 607730 DNA damage-inducible transcript 4-like protein (HIF-1 responsive protein RTP801-like) (Protein regulated in development and DNA damage response 2) (REDD-2) [REDD2] [RTP801L]
DDX60         4q32.3          Q8IY21     DDX60_HUMAN 613974 Probable ATP-dependent RNA helicase DDX60 (EC (DEAD box protein 60)
DDX60L        4q32.3          Q5H9U9     DDX6L_HUMAN        Probable ATP-dependent RNA helicase DDX60-like (EC (DEAD box protein 60-like)
DEFB131A      4p16            P59861     D131A_HUMAN        Beta-defensin 131A precursor (Beta-defensin 31) (DEFB-31) (Defensin, beta 131) [DEFB131] [DEFB31]
DGKQ          4p16.3          P52824     DGKQ_HUMAN  601207 Diacylglycerol kinase theta (EC (DAG kinase theta) (DGKtheta) (Diglyceride kinase theta) (DGK-theta) [DAGK4]
DHX15         4p15.3          O43143     DHX15_HUMAN 603403 Pre-mRNA-splicing factor ATP-dependent RNA helicase DHX15 (EC (ATP-dependent RNA helicase #46) (DEAH box protein 15) [DBP1] [DDX15]
DKK2          4q25            Q9UBU2     DKK2_HUMAN  605415 Dickkopf-related protein 2 precursor (Dickkopf-2) (Dkk-2) (hDkk-2) [UNQ682/PRO1316]
DMP1          4q21            Q13316     DMP1_HUMAN  600980 Dentin matrix acidic phosphoprotein 1 precursor (DMP-1) (Dentin matrix protein 1)
DNAJB14       4q23            Q8TBM8     DJB14_HUMAN 617487 DnaJ homolog subfamily B member 14 [UNQ9427/PRO34683]
DOK7          4p16.2          Q18PE1     DOK7_HUMAN  610285 Protein Dok-7 (Downstream of tyrosine kinase 7) [C4orf25]
DRD5          4p16.1          P21918     DRD5_HUMAN  126453 D(1B) dopamine receptor (D(5) dopamine receptor) (D1beta dopamine receptor) (Dopamine D5 receptor) [DRD1B] [DRD1L2]
DSPP          4q21.3          Q9NZW4     DSPP_HUMAN  125485 Dentin sialophosphoprotein precursor [Contains: Dentin phosphoprotein (Dentin phosphophoryn) (DPP); Dentin sialoprotein (DSP)]
DTHD1         4p14            Q6ZMT9     DTHD1_HUMAN 616979 Death domain-containing protein 1
DUX4          4q35.2          Q9UBX2     DUX4_HUMAN  606009 Double homeobox protein 4 (Double homeobox protein 10) [DUX10]
DUX4L2        4q35.2          P0CJ85     DU4L2_HUMAN        Double homeobox protein 4-like protein 2
DUX4L3        4q35.2          P0CJ86     DU4L3_HUMAN        Double homeobox protein 4-like protein 3
DUX4L4        4q35.2          P0CJ87     DU4L4_HUMAN        Double homeobox protein 4-like protein 4
DUX4L5        4q35.2          P0CJ88     DU4L5_HUMAN        Double homeobox protein 4-like protein 5
DUX4L6        4q35.2          P0CJ89     DU4L6_HUMAN        Double homeobox protein 4-like protein 6
DUX4L7        4q35.2          P0CJ90     DU4L7_HUMAN        Double homeobox protein 4-like protein 7
DUX4L9        4q35.2          Q6RFH8     DUX4C_HUMAN 615581 Double homeobox protein 4C (Double homeobox protein 4, centromeric) (DUX4c) (Double homeobox protein 4-like protein 9) [DUX4C]
EDNRA         4q31.22         P25101     EDNRA_HUMAN 131243 Endothelin-1 receptor precursor (Endothelin receptor type A) (ET-A) (ETA-R) (hET-AR) [ETA] [ETRA]
EGF           4q25            P01133     EGF_HUMAN   131530 Pro-epidermal growth factor precursor (EGF) [Contains: Epidermal growth factor (Urogastrone)]
EIF4E         4q21-q25        P06730     IF4E_HUMAN  133440 Eukaryotic translation initiation factor 4E (eIF-4E) (eIF4E) (eIF-4F 25 kDa subunit) (mRNA cap-binding protein) [EIF4EL1] [EIF4F]
ELF2          4q28            Q15723     ELF2_HUMAN         ETS-related transcription factor Elf-2 (E74-like factor 2) (New ETS-related factor) [NERF]
ELMOD2        4q31.1          Q8IZ81     ELMD2_HUMAN 610196 ELMO domain-containing protein 2
ELOVL6        4q25            Q9H5J4     ELOV6_HUMAN 611546 Elongation of very long chain fatty acids protein 6 (EC (3-keto acyl-CoA synthase ELOVL6) (ELOVL fatty acid elongase 6) (ELOVL FA elongase 6) (Fatty acid elongase 2) (hELO2) (Fatty acyl-CoA elongase) (Long-chain fatty-acyl elongase) (Very long chain 3-ketoacyl-CoA synthase 6) (Very long chain 3-oxoacyl-CoA synthase 6) [FACE] [LCE]
EMCN          4q23            Q9ULC0     MUCEN_HUMAN 608350 Endomucin precursor (Endomucin-2) (Gastric cancer antigen Ga34) (Mucin-14) (MUC-14) [EMCN2] [MUC14]
ENAM          4q13.1-q21.23   Q9NRM1     ENAM_HUMAN  606585 Enamelin precursor
ENOPH1        4q21.22         Q9UHY7     ENOPH_HUMAN        Enolase-phosphatase E1 (EC (2,3-diketo-5-methylthio-1-phosphopentane phosphatase) (MASA homolog) [MASA] [MSTP145]
ENPEP         4q25            Q07075     AMPE_HUMAN  138297 Glutamyl aminopeptidase (EC (EAP) (Aminopeptidase A) (AP-A) (Differentiation antigen gp160) (CD249 antigen)
ENPP6         4q35.1          Q6UWR7     ENPP6_HUMAN 616983 Glycerophosphocholine cholinephosphodiesterase ENPP6 precursor (EC 3.1.4.-) (EC (GPC-Cpde) (Choline-specific glycerophosphodiester phosphodiesterase) (Ectonucleotide pyrophosphatase/phosphodiesterase family member 6) (E-NPP 6) (NPP-6) [UNQ1889/PRO4334]
EPGN          4q13.3          Q6UW88     EPGN_HUMAN  618717 Epigen precursor (Epithelial mitogen) (EPG) [UNQ3072/PRO9904]
EPHA5         4q13.2          P54756     EPHA5_HUMAN 600004 Ephrin type-A receptor 5 precursor (EC (Brain-specific kinase) (EPH homology kinase 1) (EHK-1) (EPH-like kinase 7) (EK7) (hEK7) [BSK] [EHK1] [HEK7] [TYRO4]
EREG          4q13.3          O14944     EREG_HUMAN  602061 Proepiregulin precursor [Contains: Epiregulin (EPR)]
ERVMER34-1    4q12            Q9H9K5     MER34_HUMAN        Endogenous retroviral envelope protein HEMO precursor (Endogenous retrovirus group MER34 member 1 Env polyprotein) (HERV-MER_4q12 provirus ancestral Env polyprotein) (Human endogenous MER34 (medium-reiteration-frequency-family-34) open reading frame) (Human endogenous MER34 ORF) (HEMO) [Contains: Endogenous retroviral envelope protein HEMO, secreted form (Endogenous retroviral envelope protein HEMO, 48 kDa form)] [HEMO] [LP9056]
ETFDH         4q32-q35        Q16134     ETFD_HUMAN  231675 Electron transfer flavoprotein-ubiquinone oxidoreductase, mitochondrial precursor (EC (ETF-QO) (ETF-ubiquinone oxidoreductase) (Electron-transferring-flavoprotein dehydrogenase) (ETF dehydrogenase)
ETNPPL        4q25            Q8TBG4     AT2L1_HUMAN 614682 Ethanolamine-phosphate phospho-lyase (EC (Alanine--glyoxylate aminotransferase 2-like 1) [AGXT2L1]
EVC           4p16            P57679     EVC_HUMAN   604831 Ellis-van Creveld syndrome protein (DWF-1)
EVC2          4p16.1-p16.2    Q86UK5     LBN_HUMAN   607261 Limbin precursor (Ellis-van Creveld syndrome protein 2) (EVC2) [LBN]
EXOC1         4q12            Q9NV70     EXOC1_HUMAN 607879 Exocyst complex component 1 (Exocyst complex component Sec3) [SEC3] [SEC3L1] [BM-012]
EXOC1L        4q12            A0A1B0GW35 EXC1L_HUMAN        Exocyst complex component 1-like
EXOSC9        4q27            Q06265     EXOS9_HUMAN 606180 Exosome complex component RRP45 (Autoantigen PM/Scl 1) (Exosome component 9) (P75 polymyositis-scleroderma overlap syndrome-associated autoantigen) (Polymyositis/scleroderma autoantigen 1) (Polymyositis/scleroderma autoantigen 75 kDa) (PM/Scl-75) [PMSCL1]
F11           4q35            P03951     FA11_HUMAN  264900 Coagulation factor XI precursor (EC (FXI) (Plasma thromboplastin antecedent) (PTA) [Contains: Coagulation factor XIa heavy chain; Coagulation factor XIa light chain]
FABP2         4q28-q31        P12104     FABPI_HUMAN 134640 Fatty acid-binding protein, intestinal (Fatty acid-binding protein 2) (Intestinal-type fatty acid-binding protein) (I-FABP) [FABPI]
FAM114A1      4p14            Q8IWE2     NXP20_HUMAN        Protein NOXP20 (Nervous system overexpressed protein 20) (Protein FAM114A1) [NOXP20]
FAM13A        4q22.1          O94988     FA13A_HUMAN 613299 Protein FAM13A [FAM13A1] [KIAA0914]
FAM149A       4q35.2          A5PLN7     F149A_HUMAN        Protein FAM149A [MST119]
FAM184B       4p15.32         Q9ULE4     F184B_HUMAN        Protein FAM184B [KIAA1276]
FAM193A       4p16.3          P78312     F193A_HUMAN        Protein FAM193A (Protein IT14) [C4orf8] [RES4-22]
FAM200B       4p15.32         P0CF97     F200B_HUMAN        Protein FAM200B [C4orf53]
FAM218A       4q32.3          Q96MZ4     F218A_HUMAN        Protein FAM218A [C4orf39]
FAM241A       4q25            Q8N8J7     F241A_HUMAN        Uncharacterized protein FAM241A [C4orf32]
FAM47E        4q21.1          Q6ZV65     FA47E_HUMAN        Protein FAM47E
FAM53A        4p16.3          Q6NSI3     FA53A_HUMAN        Protein FAM53A (Dorsal neural-tube nuclear protein)
FAM90A26      4p16.1          D6RGX4     F90AP_HUMAN        Putative protein FAM90A26 [FAM90A26P]
FAT1          4q34-q35        Q14517     FAT1_HUMAN  600976 Protocadherin Fat 1 precursor (Cadherin family member 7) (Cadherin-related tumor suppressor homolog) (Protein fat homolog) [Contains: Protocadherin Fat 1, nuclear form] [CDHF7] [FAT]
FAT4          4q28.1          Q6V0I7     FAT4_HUMAN  612411 Protocadherin Fat 4 precursor (hFat4) (Cadherin family member 14) (FAT tumor suppressor homolog 4) (Fat-like cadherin protein FAT-J) [CDHF14] [FATJ] [Nbla00548]
FBXL5         4p15.33         Q9UKA1     FBXL5_HUMAN 605655 F-box/LRR-repeat protein 5 (F-box and leucine-rich repeat protein 5) (F-box protein FBL4/FBL5) (p45SKP2-like protein) [FBL4] [FBL5] [FLR1]
FBXO8         4q34.1          Q9NRD0     FBX8_HUMAN  605649 F-box only protein 8 (F-box/SEC7 protein FBS) [FBS] [FBX8] [DC10] [UNQ1877/PRO4320]
FBXW7         4q31.3          Q969H0     FBXW7_HUMAN 606278 F-box/WD repeat-containing protein 7 (Archipelago homolog) (hAgo) (F-box and WD-40 domain-containing protein 7) (F-box protein FBX30) (SEL-10) (hCdc4) [FBW7] [FBX30] [SEL10]
FBXW7-AS1     4q31.3          B0L3A2     DEAR_HUMAN         Dual endothelin-1(VEGFsp)/angiotensin II receptor (Dual-endothelin1/VEGFsp receptor) (DEspR protein) (FBXW7 antisense RNA 1) [DEAR] [DEspR]
FDCSP         4q13            Q8NFU4     FDSCP_HUMAN 607241 Follicular dendritic cell secreted peptide precursor (FDC secreted protein) (FDC-SP) [C4orf7] [UNQ733/PRO1419]
FGA           4q28            P02671     FIBA_HUMAN  134820 Fibrinogen alpha chain precursor [Contains: Fibrinopeptide A; Fibrinogen alpha chain]
FGB           4q28            P02675     FIBB_HUMAN  134830 Fibrinogen beta chain precursor [Contains: Fibrinopeptide B; Fibrinogen beta chain]
FGF2          4q26            P09038     FGF2_HUMAN  134920 Fibroblast growth factor 2 precursor (FGF-2) (Basic fibroblast growth factor) (bFGF) (Heparin-binding growth factor 2) (HBGF-2) [FGFB]
FGF5          4q21            P12034     FGF5_HUMAN  165190 Fibroblast growth factor 5 precursor (FGF-5) (Heparin-binding growth factor 5) (HBGF-5) (Smag-82)
FGFBP1        4p15.32         Q14512     FGFP1_HUMAN 607737 Fibroblast growth factor-binding protein 1 precursor (FGF-BP) (FGF-BP1) (FGF-binding protein 1) (FGFBP-1) (17 kDa heparin-binding growth factor-binding protein) (17 kDa HBGF-binding protein) (HBp17) [FGFBP] [HBP17]
FGFBP2        4p15.32         Q9BYJ0     FGFP2_HUMAN 607713 Fibroblast growth factor-binding protein 2 precursor (FGF-BP2) (FGF-binding protein 2) (FGFBP-2) (37 kDa killer-specific secretory protein) (Ksp37) (HBp17-related protein) (HBp17-RP) [KSP37] [UNQ425/PRO1065]
FGFR3         4p16.3          P22607     FGFR3_HUMAN 134934 Fibroblast growth factor receptor 3 precursor (EC (FGFR-3) (CD333 antigen) [JTK4]
FGFRL1        4p16            Q8N441     FGRL1_HUMAN 605830 Fibroblast growth factor receptor-like 1 precursor (FGF receptor-like protein 1) (FGF homologous factor receptor) (FGFR-like protein) (Fibroblast growth factor receptor 5) (FGFR-5) [FGFR5] [FHFR] [UNQ480/PRO943]
FGG           4q28            P02679     FIBG_HUMAN  134850 Fibrinogen gamma chain precursor [PRO2061]
FHDC1         4q31.3          Q9C0D6     FHDC1_HUMAN        FH2 domain-containing protein 1 (Inverted formin-1) [INF1] [KIAA1727]
FHIP1A        4q31.3          Q05DH4     FHI1A_HUMAN        FHF complex subunit HOOK interacting protein 1A (FHIP1A) (FTS and Hook-interacting protein like) (FHIP-L) [FAM160A1]
FIP1L1        4q12            Q6UN15     FIP1_HUMAN  607686 Pre-mRNA 3'-end-processing factor FIP1 (hFip1) (FIP1-like 1 protein) (Factor interacting with PAP) (Rearranged in hypereosinophilia) [FIP1] [RHE]
FNIP2         4q32.1          Q9P278     FNIP2_HUMAN 612768 Folliculin-interacting protein 2 (FNIP1-like protein) (O6-methylguanine-induced apoptosis 1 protein) [FNIPL] [KIAA1450] [MAPO1]
FRAS1         4q21.21         Q86XX4     FRAS1_HUMAN 607830 Extracellular matrix organizing protein FRAS1 precursor (Fraser syndrome 1 protein) [KIAA1500]
FREM3         4q31.21         P0C091     FREM3_HUMAN 608946 FRAS1-related extracellular matrix protein 3 precursor
FRG1          4q35            Q14331     FRG1_HUMAN  601278 Protein FRG1 (FSHD region gene 1 protein)
FRG2          4q35            Q64ET8     FRG2_HUMAN  609032 Protein FRG2 (FSHD region gene 2 protein) [FRG2A]
FRYL          4p12            O94915     FRYL_HUMAN         Protein furry homolog-like (ALL1-fused gene from chromosome 4p12 protein) [AF4P12] [KIAA0826]
FSTL5         4q32.2-q32.3    Q8N475     FSTL5_HUMAN        Follistatin-related protein 5 precursor (Follistatin-like protein 5) [KIAA1263]
G3BP2         4q21.1          Q9UN86     G3BP2_HUMAN        Ras GTPase-activating protein-binding protein 2 (G3BP-2) (GAP SH3 domain-binding protein 2) [KIAA0660]
GAB1          4q31.21         Q13480     GAB1_HUMAN  604439 GRB2-associated-binding protein 1 (GRB2-associated binder 1) (Growth factor receptor bound protein 2-associated protein 1)
GABRA2        4p12            P47869     GBRA2_HUMAN 137140 Gamma-aminobutyric acid receptor subunit alpha-2 precursor (GABA(A) receptor subunit alpha-2)
GABRA4        4p12            P48169     GBRA4_HUMAN 137141 Gamma-aminobutyric acid receptor subunit alpha-4 precursor (GABA(A) receptor subunit alpha-4)
GABRB1        4p12            P18505     GBRB1_HUMAN 137190 Gamma-aminobutyric acid receptor subunit beta-1 precursor (GABA(A) receptor subunit beta-1)
GABRG1        4p12            Q8N1C3     GBRG1_HUMAN 137166 Gamma-aminobutyric acid receptor subunit gamma-1 precursor (GABA(A) receptor subunit gamma-1)
GAK           4p16            O14976     GAK_HUMAN   602052 Cyclin-G-associated kinase (EC
GALNT7        4q31.1          Q86SF2     GALT7_HUMAN 605005 N-acetylgalactosaminyltransferase 7 (EC (Polypeptide GalNAc transferase 7) (GalNAc-T7) (pp-GaNTase 7) (Protein-UDP acetylgalactosaminyltransferase 7) (UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase 7)
GALNTL6       4q34.1          Q49A17     GLTL6_HUMAN 615138 Polypeptide N-acetylgalactosaminyltransferase-like 6 (EC (Polypeptide GalNAc transferase 17) (GalNAc-T17) (pp-GaNTase 17) (Protein-UDP acetylgalactosaminyltransferase 17) (Putative polypeptide N-acetylgalactosaminyltransferase 17) (UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase 17) [GALNT17]
GAR1          4q25            Q9NY12     GAR1_HUMAN  606468 H/ACA ribonucleoprotein complex subunit 1 (Nucleolar protein family A member 1) (snoRNP protein GAR1) [NOLA1]
GASK1B        4q32.1          Q6UWH4     GAK1B_HUMAN        Golgi-associated kinase 1B (Expressed in nerve and epithelium during development) (Protein FAM198B) [C4orf18] [ENED] [FAM198B] [AD021] [UNQ2512/PRO6001]
GATB          4q27-q28        O75879     GATB_HUMAN  603645 Glutamyl-tRNA(Gln) amidotransferase subunit B, mitochondrial precursor (EC 6.3.5.-) (Glu-AdT subunit B) (Cytochrome c oxidase assembly factor PET112 homolog) [PET112] [PET112L] [HSPC199]
GBA3          4p15.31         Q9H227     GBA3_HUMAN  606619 Cytosolic beta-glucosidase (EC (Cytosolic beta-glucosidase-like protein 1) (Cytosolic glycosylceramidase) (Cytosolic GCase) (Glucosidase beta acid 3) (Glucosylceramidase beta 3) (Klotho-related protein) (KLrP) [CBG] [CBGL1]
GC            4q12-q13        P02774     VTDB_HUMAN  139200 Vitamin D-binding protein precursor (DBP) (VDB) (Gc protein-derived macrophage activating factor) (Gc-MAF) (GcMAF) (Gc-globulin) (Group-specific component) (Gc) (Vitamin D-binding protein-macrophage activating factor) (DBP-maf)
GIMD1         4q24            P0DJR0     GIMD1_HUMAN        GTPase IMAP family member GIMD1 (GIMAP family P-loop NTPase domain-containing protein 1)
GK2           4q13            Q14410     GLPK2_HUMAN 600148 Glycerol kinase 2 (EC (GK 2) (Glycerokinase 2) (ATP:glycerol 3-phosphotransferase 2) (Glycerol kinase, testis specific 2) [GKP2] [GKTA]
GK3P          4q32            Q14409     GLPK3_HUMAN 600149 Glycerol kinase 3 (EC (GK 3) (Glycerokinase 3) (ATP:glycerol 3-phosphotransferase 3) (Glycerol kinase 3 pseudogene) (Glycerol kinase, testis specific 1) [GKP3] [GKTB]
GLRA3         4q33-q34        O75311     GLRA3_HUMAN 600421 Glycine receptor subunit alpha-3 precursor
GLRB          4q31.3          P48167     GLRB_HUMAN  138492 Glycine receptor subunit beta precursor (Glycine receptor 58 kDa subunit)
GNPDA2        4p13            Q8TDQ7     GNPI2_HUMAN 613222 Glucosamine-6-phosphate isomerase 2 (EC (Glucosamine-6-phosphate deaminase 2) (GNPDA 2) (GlcN6P deaminase 2) (Glucosamine-6-phosphate isomerase SB52) [GNP2]
GNRHR         4q21.2          P30968     GNRHR_HUMAN 138850 Gonadotropin-releasing hormone receptor (GnRH receptor) (GnRH-R) [GRHR]
GPAT3         4q21.23         Q53EU6     GPAT3_HUMAN 610958 Glycerol-3-phosphate acyltransferase 3 (EC (GPAT-3) (1-acyl-sn-glycerol-3-phosphate O-acyltransferase 10) (AGPAT 10) (1-acyl-sn-glycerol-3-phosphate O-acyltransferase 9) (EC (1-AGP acyltransferase 9) (1-AGPAT 9) (Acyl-CoA:glycerol-3-phosphate acyltransferase 3) (hGPAT3) (Lung cancer metastasis-associated protein 1) (Lysophosphatidic acid acyltransferase theta) (LPAAT-theta) (MAG-1) [AGPAT9] [MAG1] [HMFN0839] [UNQ2753/PRO6492]
GPM6A         4q34            P51674     GPM6A_HUMAN 601275 Neuronal membrane glycoprotein M6-a (M6a) [M6A]
GPR78         4p16.1          Q96P69     GPR78_HUMAN 606921 G-protein coupled receptor 78 [UNQ5925/PRO19818]
GPRIN3        4q22.1          Q6ZVF9     GRIN3_HUMAN 611241 G protein-regulated inducer of neurite outgrowth 3 (GRIN3) [KIAA2027]
GRIA2         4q32-q33        P42262     GRIA2_HUMAN 138247 Glutamate receptor 2 precursor (GluR-2) (AMPA-selective glutamate receptor 2) (GluR-B) (GluR-K2) (Glutamate receptor ionotropic, AMPA 2) (GluA2) [GLUR2]
GRID2         4q22            O43424     GRID2_HUMAN 602368 Glutamate receptor ionotropic, delta-2 precursor (GluD2) (GluR delta-2 subunit) [GLURD2]
GRK4          4p16.3          P32298     GRK4_HUMAN  137026 G protein-coupled receptor kinase 4 (EC (G protein-coupled receptor kinase GRK4) (ITI1) [GPRK2L] [GPRK4]
GRPEL1        4p16            Q9HAV7     GRPE1_HUMAN 606173 GrpE protein homolog 1, mitochondrial precursor (HMGE) (Mt-GrpE#1) [GREPEL1]
GRSF1         4q13            Q12849     GRSF1_HUMAN 604851 G-rich sequence factor 1 precursor (GRSF-1)
GRXCR1        4p13            A8MXD5     GRCR1_HUMAN 613283 Glutaredoxin domain-containing cysteine-rich protein 1 [DFNB25]
GSTCD         4q24            Q8NEC7     GSTCD_HUMAN 615912 Glutathione S-transferase C-terminal domain-containing protein
GSX2          4q11-q12        Q9BZM3     GSX2_HUMAN  616253 GS homeobox 2 (Genetic-screened homeobox 2) (Homeobox protein GSH-2) [GSH2]
GUCY1A1       4q31.3-q33      Q02108     GCYA1_HUMAN 139396 Guanylate cyclase soluble subunit alpha-1 (EC (GCS-alpha-1) (Guanylate cyclase soluble subunit alpha-3) (GCS-alpha-3) (Soluble guanylate cyclase large subunit) [GUC1A3] [GUCSA3] [GUCY1A3]
GUCY1B1       4q31.3-q33      Q02153     GCYB1_HUMAN 139397 Guanylate cyclase soluble subunit beta-1 (EC (GCS-beta-1) (Guanylate cyclase soluble subunit beta-3) (GCS-beta-3) (Soluble guanylate cyclase small subunit) [GUC1B3] [GUCSB3] [GUCY1B3]
GUF1          4p13            Q8N442     GUF1_HUMAN  617064 Translation factor GUF1, mitochondrial precursor (EC 3.6.5.-) (Elongation factor 4 homolog) (EF-4) (GTPase GUF1) (Ribosomal back-translocase)
GYPA          4q28.2-q31.1    P02724     GLPA_HUMAN  617922 Glycophorin-A precursor (MN sialoglycoprotein) (PAS-2) (Sialoglycoprotein alpha) (CD235a antigen) [GPA]
GYPB          4q28-q31        P06028     GLPB_HUMAN  617923 Glycophorin-B precursor (PAS-3) (SS-active sialoglycoprotein) (Sialoglycoprotein delta) (CD235b antigen) [GPB]
GYPE          4q31.1          P15421     GLPE_HUMAN  138590 Glycophorin-E precursor [GPE]
H2AZ1         4q24            P0C0S5     H2AZ_HUMAN  142763 Histone H2A.Z (H2A/z) [H2AFZ] [H2AZ]
HADH          4q22-q26        Q16836     HCDH_HUMAN  601609 Hydroxyacyl-coenzyme A dehydrogenase, mitochondrial precursor (EC (HCDH) (Medium and short-chain L-3-hydroxyacyl-coenzyme A dehydrogenase) (Short-chain 3-hydroxyacyl-CoA dehydrogenase) [HAD] [HAD1] [HADHSC] [SCHAD]
HAND2         4q33            P61296     HAND2_HUMAN 602407 Heart- and neural crest derivatives-expressed protein 2 (Class A basic helix-loop-helix protein 26) (bHLHa26) (Deciduum, heart, autonomic nervous system and neural crest derivatives-expressed protein 2) (dHAND) [BHLHA26] [DHAND]
HAUS3         4p16.3          Q68CZ6     HAUS3_HUMAN 613430 HAUS augmin-like complex subunit 3 [C4orf15]
HELQ          4q21.23         Q8TDG4     HELQ_HUMAN  606769 Helicase POLQ-like (EC (Mus308-like helicase) (POLQ-like helicase) [HEL308]
HELT          4q35.1          A6NFD8     HELT_HUMAN  617546 Hairy and enhancer of split-related protein HELT (HES/HEY-like transcription factor)
HERC3         4q21            Q15034     HERC3_HUMAN 605200 Probable E3 ubiquitin-protein ligase HERC3 (EC (HECT domain and RCC1-like domain-containing protein 3) (HECT-type E3 ubiquitin transferase HERC3) [KIAA0032]
HERC5         4q22.1          Q9UII4     HERC5_HUMAN 608242 E3 ISG15--protein ligase HERC5 (EC 2.3.2.-) (Cyclin-E-binding protein 1) (HECT domain and RCC1-like domain-containing protein 5) [CEB1] [CEBP1]
HERC6         4q22            Q8IVU3     HERC6_HUMAN 609249 Probable E3 ubiquitin-protein ligase HERC6 (EC (HECT domain and RCC1-like domain-containing protein 6) (HECT-type E3 ubiquitin transferase HERC6)
HGFAC         4p16            Q04756     HGFA_HUMAN  604552 Hepatocyte growth factor activator precursor (EC 3.4.21.-) (HGF activator) (HGFA) [Contains: Hepatocyte growth factor activator short chain; Hepatocyte growth factor activator long chain]
HHIP          4q28-q32        Q96QV1     HHIP_HUMAN  606178 Hedgehog-interacting protein precursor (HHIP) (HIP) [HIP] [UNQ5825/PRO19644]
HMGB2         4q31            P26583     HMGB2_HUMAN 163906 High mobility group protein B2 (High mobility group protein 2) (HMG-2) [HMG2]
HMX1          4p16.1          Q9NP08     HMX1_HUMAN  142992 Homeobox protein HMX1 (Homeobox protein H6) [H6]
HNRNPD        4q21.1-q21.2    Q14103     HNRPD_HUMAN 601324 Heterogeneous nuclear ribonucleoprotein D0 (hnRNP D0) (AU-rich element RNA-binding protein 1) [AUF1] [HNRPD]
HNRNPDL       4q21.22         O14979     HNRDL_HUMAN 607137 Heterogeneous nuclear ribonucleoprotein D-like (hnRNP D-like) (hnRNP DL) (AU-rich element RNA-binding factor) (JKT41-binding protein) (Protein laAUF1) [HNRPDL] [JKTBP]
HOPX          4q11-q12        Q9BPY8     HOP_HUMAN   607275 Homeodomain-only protein (Lung cancer-associated Y protein) (Not expressed in choriocarcinoma protein 1) (Odd homeobox protein 1) [HOD] [HOP] [LAGY] [NECC1] [OB1]
HPF1          4q33            Q9NWY4     HPF1_HUMAN  616614 Histone PARylation factor 1 [C4orf27]
HPGD          4q34-q35        P15428     PGDH_HUMAN  601688 15-hydroxyprostaglandin dehydrogenase [NAD(+)] (EC (15-PGDH) (Prostaglandin dehydrogenase 1) (Short chain dehydrogenase/reductase family 36C member 1) [PGDH1] [SDR36C1]
HPGDS         4q22.1          O60760     HPGDS_HUMAN 602598 Hematopoietic prostaglandin D synthase (EC (H-PGDS) (GST class-sigma) (Glutathione S-transferase) (EC (Glutathione-dependent PGD synthase) (Glutathione-requiring prostaglandin D synthase) (Prostaglandin-H2 D-isomerase) [GSTS] [PGDS] [PTGDS2]
HPSE          4q21.3          Q9Y251     HPSE_HUMAN  604724 Heparanase precursor (EC (Endo-glucoronidase) (Heparanase-1) (Hpa1) [Contains: Heparanase 8 kDa subunit; Heparanase 50 kDa subunit] [HEP] [HPA] [HPA1] [HPR1] [HPSE1] [HSE1]
HS3ST1        4p16            O14792     HS3S1_HUMAN 603244 Heparan sulfate glucosamine 3-O-sulfotransferase 1 precursor (EC (Heparan sulfate D-glucosaminyl 3-O-sulfotransferase 1) (3-OST-1) (Heparan sulfate 3-O-sulfotransferase 1) (h3-OST-1) [3OST] [3OST1]
HSD17B11      4q22.1          Q8NBQ5     DHB11_HUMAN 612831 Estradiol 17-beta-dehydrogenase 11 precursor (EC (17-beta-hydroxysteroid dehydrogenase 11) (17-beta-HSD 11) (17bHSD11) (17betaHSD11) (17-beta-hydroxysteroid dehydrogenase XI) (17-beta-HSD XI) (17betaHSDXI) (Cutaneous T-cell lymphoma-associated antigen HD-CL-03) (CTCL-associated antigen HD-CL-03) (Dehydrogenase/reductase SDR family member 8) (Retinal short-chain dehydrogenase/reductase 2) (retSDR2) (Short chain dehydrogenase/reductase family 16C member 2) [DHRS8] [PAN1B] [SDR16C2] [PSEC0029] [UNQ207/PRO233]
HSD17B13      4q22.1          Q7Z5P4     DHB13_HUMAN 612127 17-beta-hydroxysteroid dehydrogenase 13 precursor (EC 1.1.-.-) (17-beta-HSD 13) (Short chain dehydrogenase/reductase family 16C member 3) (Short-chain dehydrogenase/reductase 9) [SCDR9] [SDR16C3] [HMFN0376] [UNQ497/PRO1014]
HSP90AA4P     4q35.2          Q58FG1     HS904_HUMAN        Putative heat shock protein HSP 90-alpha A4 (Heat shock 90 kDa protein 1 alpha-like 2) (Heat shock protein 90-alpha D) (Heat shock protein 90Ad) [HSP90AD] [HSPCAL2]
HSP90AB2P     4p15.33         Q58FF8     H90B2_HUMAN        Putative heat shock protein HSP 90-beta 2 (Heat shock protein 90-beta b) (Heat shock protein 90Bb) [HSP90BB]
HSP90AB3P     4q21-q25        Q58FF7     H90B3_HUMAN        Putative heat shock protein HSP 90-beta-3 (Heat shock protein 90-beta c) (Heat shock protein 90Bc) [HSP90BC]
HSPA4L        4q28            O95757     HS74L_HUMAN 619077 Heat shock 70 kDa protein 4L (Heat shock 70-related protein APG-1) (Heat-shock protein family A member 4-like protein) (HSPA4-like protein) (Osmotic stress protein 94) [APG1] [OSP94]
HTN1          4q12-q21        P15515     HIS1_HUMAN  142701 Histatin-1 precursor (Histidine-rich protein 1) (Post-PB protein) (PPB) [Contains: His1-(31-57)-peptide (His1 31/57) (His1-(12-38)-peptide) (His1 12/38) (Histatin-2)] [HIS1]
HTN3          4q12-q21        P15516     HIS3_HUMAN  142702 Histatin-3 precursor (Basic histidine-rich protein) (Hst) (Histidine-rich protein 3) (PB) [Contains: Histatin-3; His3-(20-44)-peptide (His3 20/44) (His3-(1-25)-peptide) (His3 1/25) (Histatin-3 1/25) (Histatin-6); His3-(20-43)-peptide (His3 20/43) (His3-(1-24)-peptide) (His3 1/24) (Histatin-3 1/24) (Histatin-5); His3-(20-32)-peptide (His3 20/32) (His3-(1-13)-peptide) (His3 1/13) (Histatin-3 1/13); His3-(20-31)-peptide (His3 20/31) (His3-(1-12)-peptide) (His3 1/12) (Histatin-3 1/12); His3-(20-30)-peptide (His3 20/30) (His3-(1-11)-peptide) (His3 1/11) (Histatin-3 1/11); His3-(24-32)-peptide (His3 24/32) (His3-(5-13)-peptide) (His3 5/13) (Histatin-3 5/13); His3-(24-31)-peptide (His3 24/31) (His3-(5-12)-peptide) (His3 5/12) (Histatin-11) (Histatin-3 5/12); His3-(24-30)-peptide (His3 24/30) (His3-(5-11)-peptide) (His3 5/11) (Histatin-12) (Histatin-3 5/11); His3-(25-32)-peptide (His3 25/32) (His3-(6-13)-peptide) (His3 6/13) (Histatin-3 6/13); His3-(25-30)-peptide (His3 25/30) (His3-(6-11)-peptide) (His3 6/11) (Histatin-3 6/11); His3-(26-32)-peptide (His3 26/32) (His3-(7-13)-peptide) (His3 7/13) (Histatin-3 7/13); His3-(26-31)-peptide (His3 26/31) (His3-(7-12)-peptide) (His3 7/12) (Histatin-3 7/12); His3-(26-30)-peptide (His3 26/30) (His3-(7-11)-peptide) (His3 7/11) (Histatin-3 7/11); His3-(31-51)-peptide (His3 31/51) (His3-(12-32)-peptide) (His3 12/32) (Histatin-3 12/32) (Histatin-4); His3-(31-44)-peptide (His3 31/44) (His3-(12-25)-peptide) (His3 12/25) (Histatin-3 12/25) (Histatin-9); His3-(31-43)-peptide (His3 31/43) (His3-(12-24)-peptide) (His3 12/24) (Histatin-3 12/24) (Histatin-7); His3-(32-44)-peptide (His3 32/44) (His3-(13-25)-peptide) (His3 13/25) (Histatin-10) (Histatin-3 13/25); His3-(32-43)-peptide (His3 32-43) (His3-(13-24)-peptide) (His3 13/24) (Histatin-3 13/24) (Histatin-8); His3-(33-44)-peptide (His3 33/44) (His3-(14-25)-peptide) (His3 14/25) (Histatin-3 14/25); His3-(33-43)-peptide (His3 33/43) (His3-(14-24)-peptide) (His3 14/24) (Histatin-3 14/24); His3-(34-44)-peptide (His3 34/44) (His3-(15-25)-peptide) (His3 15/25) (Histatin-3 15/25); His3-(34-43)-peptide (His3 34/43) (His3-(15-24)-peptide) (His3 15/24) (Histatin-3 15/24); His3-(45-51)-peptide (His3 45/51) (His3-(26-32)-peptide) (His3 26/32) (Histatin-3 26/32); His3-(47-51)-peptide (His3 47/51) (His3-(28-32)-peptide) (His3 28/32) (Histatin-3 28/32); His3-(48-51)-peptide (His3 48/51) (His3-(29-32)-peptide) (His3 29/32) (Histatin-3 29/32)] [HIS2]
HTRA3         4p16.1          P83110     HTRA3_HUMAN 608785 Serine protease HTRA3 precursor (EC 3.4.21.-) (High-temperature requirement factor A3) (Pregnancy-related serine protease) [PRSP]
HTT           4p16.3          P42858     HD_HUMAN    613004 Huntingtin (Huntington disease protein) (HD protein) [Contains: Huntingtin, myristoylated N-terminal fragment] [HD] [IT15]
IBSP          4q28-q31        P21815     SIAL_HUMAN  147563 Bone sialoprotein 2 precursor (Bone sialoprotein II) (BSP II) (Cell-binding sialoprotein) (Integrin-binding sialoprotein) [BNSP]
IDUA          4p16.3          P35475     IDUA_HUMAN  252800 Alpha-L-iduronidase precursor (EC
IGFBP7        4q12            Q16270     IBP7_HUMAN  602867 Insulin-like growth factor-binding protein 7 precursor (IBP-7) (IGF-binding protein 7) (IGFBP-7) (IGFBP-rP1) (MAC25 protein) (PGI2-stimulating factor) (Prostacyclin-stimulating factor) (Tumor-derived adhesion factor) (TAF) [MAC25] [PSF]
IL15          4q31            P40933     IL15_HUMAN  600554 Interleukin-15 precursor (IL-15)
IL2           4q26-q27        P60568     IL2_HUMAN   147680 Interleukin-2 precursor (IL-2) (T-cell growth factor) (TCGF) (Aldesleukin)
IL21          4q26-q27        Q9HBE4     IL21_HUMAN  605384 Interleukin-21 precursor (IL-21) (Za11)
ING2          4q35.1          Q9H160     ING2_HUMAN  604215 Inhibitor of growth protein 2 (Inhibitor of growth 1-like protein) (ING1Lp) (p32) (p33ING2) [ING1L]
INPP4B        4q31.21         O15327     INP4B_HUMAN 607494 Inositol polyphosphate 4-phosphatase type II (Type II inositol 3,4-bisphosphate 4-phosphatase) (EC
INTS12        4q24            Q96CB8     INT12_HUMAN 611355 Integrator complex subunit 12 (Int12) (PHD finger protein 22) [PHF22] [SBBI22]
INTU          4q28.1          Q9ULD6     INTU_HUMAN  610621 Protein inturned (Inturned planar cell polarity effector homolog) (PDZ domain-containing protein 6) [KIAA1284] [PDZD6] [PDZK6]
IQCM          4q31.23         A0A1B0GVH7 IQCM_HUMAN         IQ domain-containing protein M
IRF2          4q34.1-q35.1    P14316     IRF2_HUMAN  147576 Interferon regulatory factor 2 (IRF-2)
JADE1         4q28.2          Q6IE81     JADE1_HUMAN 610514 Protein Jade-1 (Jade family PHD finger protein 1) (PHD finger protein 17) [KIAA1807] [PHF17]
JAKMIP1       4p16.1          Q96N16     JKIP1_HUMAN 611195 Janus kinase and microtubule-interacting protein 1 (GABA-B receptor-binding protein) (Multiple alpha-helices and RNA-linker protein 1) (Marlin-1) [GABABRBP] [JAMIP1] [MARLIN1]
JCHAIN        4q21            P01591     IGJ_HUMAN   147790 Immunoglobulin J chain precursor (Joining chain of multimeric IgA and IgM) [IGCJ] [IGJ]
KCNIP4        4p15.31         Q6PIL6     KCIP4_HUMAN 608182 Kv channel-interacting protein 4 (KChIP4) (A-type potassium channel modulatory protein 4) (Calsenilin-like protein) (Potassium channel-interacting protein 4) [CALP] [KCHIP4]
KCTD8         4p13            Q6ZWB6     KCTD8_HUMAN 618442 BTB/POZ domain-containing protein KCTD8
KDR           4q11-q12        P35968     VGFR2_HUMAN 191306 Vascular endothelial growth factor receptor 2 precursor (EC (VEGFR-2) (Fetal liver kinase 1) (FLK-1) (Kinase insert domain receptor) (KDR) (Protein-tyrosine kinase receptor flk-1) (CD309 antigen) [FLK1] [VEGFR2]
KIAA0232      4p16.1          Q92628     K0232_HUMAN 619237 Uncharacterized protein KIAA0232
KIAA1109      4q27            Q2LD37     K1109_HUMAN 611565 Transmembrane protein KIAA1109 (Fragile site-associated protein) [FSA] [KIAA1371]
KIT           4q11-q12        P10721     KIT_HUMAN   164920 Mast/stem cell growth factor receptor Kit precursor (EC (SCFR) (Piebald trait protein) (PBT) (Proto-oncogene c-Kit) (Tyrosine-protein kinase Kit) (p145 c-kit) (v-kit Hardy-Zuckerman 4 feline sarcoma viral oncogene homolog) (CD117 antigen) [SCFR]
KLB           4p14            Q86Z14     KLOTB_HUMAN 611135 Beta-klotho (BKL) (BetaKlotho) (Klotho beta-like protein)
KLF3          4p14            P57682     KLF3_HUMAN  609392 Krueppel-like factor 3 (Basic krueppel-like factor) (CACCC-box-binding protein BKLF) (TEF-2) [BKLF]
KLHL2         4q21.2          O95198     KLHL2_HUMAN 605774 Kelch-like protein 2 (Actin-binding protein Mayven)
KLHL5         4p15.1          Q96PQ7     KLHL5_HUMAN 608064 Kelch-like protein 5
KLHL8         4q22.1          Q9P2G9     KLHL8_HUMAN 611967 Kelch-like protein 8 [KIAA1378]
KLKB1         4q34-q35        P03952     KLKB1_HUMAN 229000 Plasma kallikrein precursor (EC (Fletcher factor) (Kininogenin) (Plasma prekallikrein) (PKK) [Contains: Plasma kallikrein heavy chain; Plasma kallikrein light chain] [KLK3]
LAMTOR3       4q23            Q9UHA4     LTOR3_HUMAN 603296 Ragulator complex protein LAMTOR3 (Late endosomal/lysosomal adaptor and MAPK and MTOR activator 3) (MEK-binding partner 1) (Mp1) (Mitogen-activated protein kinase kinase 1-interacting protein 1) (Mitogen-activated protein kinase scaffold protein 1) [MAP2K1IP1] [MAPKSP1] [PRO2783]
LAP3          4p15.32         P28838     AMPL_HUMAN  170250 Cytosol aminopeptidase (EC (Cysteinylglycine-S-conjugate dipeptidase) (EC (Leucine aminopeptidase 3) (LAP-3) (Leucyl aminopeptidase) (Peptidase S) (Proline aminopeptidase) (EC (Prolyl aminopeptidase) [LAPEP] [PEPS]
LARP1B        4q28.2          Q659C4     LAR1B_HUMAN        La-related protein 1B (La ribonucleoprotein domain family member 1B) (La ribonucleoprotein domain family member 2) (La-related protein 2) [LARP2]
LARP7         4q25            Q4G0J3     LARP7_HUMAN 612026 La-related protein 7 (La ribonucleoprotein domain family member 7) (hLARP7) (P-TEFb-interaction protein for 7SK stability) (PIP7S) [HDCMA18P]
LCORL         4p15.32         Q8N3X6     LCORL_HUMAN 611799 Ligand-dependent nuclear receptor corepressor-like protein (LCoR-like protein)
LDB2          4p16            O43679     LDB2_HUMAN  603450 LIM domain-binding protein 2 (LDB-2) (Carboxyl-terminal LIM domain-binding protein 1) (CLIM-1) (LIM domain-binding factor CLIM1) [CLIM1]
LEF1          4q23-q25        Q9UJU2     LEF1_HUMAN  153245 Lymphoid enhancer-binding factor 1 (LEF-1) (T cell-specific transcription factor 1-alpha) (TCF1-alpha)
LETM1         4p16.3          O95202     LETM1_HUMAN 604407 Mitochondrial proton/calcium exchanger protein precursor (Leucine zipper-EF-hand-containing transmembrane protein 1)
LGI2          4p15.2          Q8N0V4     LGI2_HUMAN  608301 Leucine-rich repeat LGI family member 2 precursor (LGI1-like protein 2) (Leucine-rich glioma-inactivated protein 2) [KIAA1916] [LGIL2]
LIAS          4p14            O43766     LIAS_HUMAN  607031 Lipoyl synthase, mitochondrial precursor (EC (Lipoate synthase) (LS) (Lip-syn) (Lipoic acid synthase) [LAS] [HUSSY-01]
LIMCH1        4p13            Q9UPQ0     LIMC1_HUMAN 617750 LIM and calponin homology domains-containing protein 1 [KIAA1102]
LIN54         4q21.22         Q6MZP7     LIN54_HUMAN 613367 Protein lin-54 homolog (CXC domain-containing protein 1) [CXCDC1] [KIAA2037]
LINC00575     4q21.3          Q6W349     CD011_HUMAN        Putative uncharacterized protein encoded by LINC00575 [C4orf11]
LINC01587     4p16.2          Q99440     CD006_HUMAN        Uncharacterized protein encoded by LINC01587 (Protein AC1) [AC1] [C4orf6]
LNX1          4q12            Q8TBB1     LNX1_HUMAN  609732 E3 ubiquitin-protein ligase LNX (EC (Ligand of Numb-protein X 1) (Numb-binding protein 1) (PDZ domain-containing RING finger protein 2) (RING-type E3 ubiquitin transferase LNX) [LNX] [PDZRN2] [UNQ574/PRO1136]
LRAT          4q31.3          O95237     LRAT_HUMAN  604863 Lecithin retinol acyltransferase (EC (Phosphatidylcholine--retinol O-acyltransferase)
LRBA          4q31.23         P50851     LRBA_HUMAN  606453 Lipopolysaccharide-responsive and beige-like anchor protein (Beige-like protein) (CDC4-like protein) [BGL] [CDC4L] [LBA]
LRIT3         4q25            Q3SXY7     LRIT3_HUMAN 615004 Leucine-rich repeat, immunoglobulin-like domain and transmembrane domain-containing protein 3 precursor
LRP2BP        4q35.1          Q9P2M1     LR2BP_HUMAN 619020 LRP2-binding protein (Megalin-binding protein) (MegBP) [KIAA1325]
LRPAP1        4p16.3          P30533     AMRP_HUMAN  104225 Alpha-2-macroglobulin receptor-associated protein precursor (Alpha-2-MRAP) (Low density lipoprotein receptor-related protein-associated protein 1) (RAP) [A2MRAP]
LRRC66        4q12            Q68CR7     LRC66_HUMAN        Leucine-rich repeat-containing protein 66
LSM6          4q31.21         P62312     LSM6_HUMAN  607286 U6 snRNA-associated Sm-like protein LSm6
LYAR          4p16.3          Q9NX58     LYAR_HUMAN  617684 Cell growth-regulating nucleolar protein [PNAS-5]
MAB21L2       4q31.3          Q9Y586     MB212_HUMAN 604357 Protein mab-21-like 2
MAD2L1        4q27            Q13257     MD2L1_HUMAN 601467 Mitotic spindle assembly checkpoint protein MAD2A (HsMAD2) (Mitotic arrest deficient 2-like protein 1) (MAD2-like protein 1) [MAD2]
MAEA          4p16.3          Q7L5Y9     MAEA_HUMAN  606801 E3 ubiquitin-protein transferase MAEA (EC (Cell proliferation-inducing gene 5 protein) (Erythroblast macrophage protein) (Human lung cancer oncogene 10 protein) (HLC-10) (Macrophage erythroblast attacher) (P44EMLP) [EMP] [HLC10] [PIG5]
MAFIP         4               Q8WZ33     MAFIP_HUMAN        MaFF-interacting protein (Tektin-4 like protein PP5644) [MIP] [PP5644]
MAML3         4q28            Q96JK9     MAML3_HUMAN 608991 Mastermind-like protein 3 (Mam-3) [KIAA1816]
MAN2B2        4p16.2          Q9Y2E5     MA2B2_HUMAN 618899 Epididymis-specific alpha-mannosidase precursor (EC (Alpha-1,6-mannosidase) (Mannosidase alpha class 2B member 2) [KIAA0935]
MANBA         4q22-q25        O00462     MANBA_HUMAN 609489 Beta-mannosidase precursor (EC (Lysosomal beta A mannosidase) (Mannanase) (Mannase) [MANB1]
MAP9          4q32.1          Q49MG5     MAP9_HUMAN  610070 Microtubule-associated protein 9 (Aster-associated protein) [ASAP]
MAPK10        4q22.1-q23      P53779     MK10_HUMAN  602897 Mitogen-activated protein kinase 10 (EC (MAP kinase 10) (MAPK 10) (MAP kinase p49 3F12) (Stress-activated protein kinase 1b) (SAPK1b) (Stress-activated protein kinase JNK3) (c-Jun N-terminal kinase 3) [JNK3] [JNK3A] [PRKM10] [SAPK1B]
MARCHF1       4q32.2          Q8TCQ1     MARH1_HUMAN 613331 E3 ubiquitin-protein ligase MARCHF1 (EC (Membrane-associated RING finger protein 1) (Membrane-associated RING-CH protein I) (MARCH-I) (RING finger protein 171) (RING-type E3 ubiquitin transferase MARCHF1) [MARCH1] [RNF171]
MCUB          4q25            Q9NWR8     MCUB_HUMAN         Calcium uniporter regulatory subunit MCUb, mitochondrial precursor (MCUb) (Coiled-coil domain-containing protein 109B) [CCDC109B]
MED28         4p16            Q9H204     MED28_HUMAN 610311 Mediator of RNA polymerase II transcription subunit 28 (Endothelial-derived protein 1) (Mediator complex subunit 28) (Merlin and Grb2-interacting cytoskeletal protein) (Magicin) (Tumor angiogenesis marker EG-1) [EG1] [FKSG20]
MEPE          4q21.1          Q9NQ76     MEPE_HUMAN  605912 Matrix extracellular phosphoglycoprotein precursor (Osteoblast/osteocyte factor 45) (OF45) (Osteoregulin)
METAP1        4q23            P53582     MAP11_HUMAN 610151 Methionine aminopeptidase 1 (EC (MAP 1) (MetAP 1) (Peptidase M 1) [KIAA0094]
METTL14       4q26            Q9HCE5     MET14_HUMAN 616504 N6-adenosine-methyltransferase non-catalytic subunit (Methyltransferase-like protein 14) (hMETTL14) [KIAA1627]
MFAP3L        4q32            O75121     MFA3L_HUMAN        Microfibrillar-associated protein 3-like precursor (Testis development protein NYD-SP9) [KIAA0626] [HSD-39] [HSD39]
MFSD10        4p16.3          Q14728     MFS10_HUMAN 610977 Major facilitator superfamily domain-containing protein 10 (Tetracycline transporter-like protein) [TETRAN]
MFSD8         4q28.1-q28.2    Q8NHS3     MFSD8_HUMAN 611124 Major facilitator superfamily domain-containing protein 8 (Ceroid-lipofuscinosis neuronal protein 7) [CLN7]
MGARP         4q31.1          Q8TDB4     HUMMR_HUMAN        Protein MGARP (Corneal endothelium-specific protein 1) (CESP-1) (Hypoxia up-regulated mitochondrial movement regulator protein) (Mitochondria-localized glutamic acid-rich protein) (Ovary-specific acidic protein) [C4orf49] [CESP1] [HUMMR] [OSAP] [GS3582]
MGAT4D        4q31            A6NG13     MGT4D_HUMAN 610310 Alpha-1,3-mannosyl-glycoprotein 4-beta-N-acetylglucosaminyltransferase-like protein MGAT4D (N-acetylglucosaminyltransferase MGAT1 inhibitory protein) (GlcNAcT-I inhibitory protein) (GnT1IP)
MGST2         4q28.3          Q99735     MGST2_HUMAN 601733 Microsomal glutathione S-transferase 2 (EC (Microsomal GST-2) (Glutathione peroxidase MGST2) (EC 1.11.1.-) (Leukotriene C4 synthase MGST2) (EC (Microsomal glutathione S-transferase II) (Microsomal GST-II) [GST2]
MMAA          4q31.21         Q8IVH4     MMAA_HUMAN  607481 Methylmalonic aciduria type A protein, mitochondrial precursor (EC 3.6.-.-)
MMRN1         4q22            Q13201     MMRN1_HUMAN 601456 Multimerin-1 precursor (EMILIN-4) (Elastin microfibril interface located protein 4) (Elastin microfibril interfacer 4) (Endothelial cell multimerin) [Contains: Platelet glycoprotein Ia*; 155 kDa platelet multimerin (p-155) (p155)] [ECM] [EMILIN4] [GPIA*] [MMRN]
MND1          4q31.3          Q9BWT6     MND1_HUMAN  611422 Meiotic nuclear division protein 1 homolog [GAJ]
MOB1B         4q13.3          Q7L9L4     MOB1B_HUMAN 609282 MOB kinase activator 1B (Mob1 homolog 1A) (Mob1A) (Mob1B) (Mps one binder kinase activator-like 1A) [MOB4A] [MOBKL1A]
MRFAP1        4p16.1          Q9Y605     MOFA1_HUMAN 616905 MORF4 family-associated protein 1 (Protein PGR1) (Protein associated with MRG of 14 kDa) [PAM14] [PGR1]
MRFAP1L1      4p16.1          Q96HT8     MR1L1_HUMAN        MORF4 family-associated protein 1-like 1 [PP784]
MRPL1         4q21.1          Q9BYD6     RM01_HUMAN  611821 39S ribosomal protein L1, mitochondrial precursor (L1mt) (MRP-L1) (Mitochondrial large ribosomal subunit protein uL1m) [BM-022]
MRPS18C       4q21.23         Q9Y3D5     RT18C_HUMAN 611983 28S ribosomal protein S18c, mitochondrial precursor (MRP-S18-c) (Mrps18-c) (S18mt-c) (28S ribosomal protein S18-1, mitochondrial) (MRP-S18-1) (Mitochondrial small ribosomal subunit protein bS18c) (Mitochondrial small ribosomal subunit protein bS18m) [CGI-134]
MSANTD1       4p16.2          Q6ZTZ1     MSD1_HUMAN         Myb/SANT-like DNA-binding domain-containing protein 1 [C4orf44]
MSMO1         4q32-q34        Q15800     MSMO1_HUMAN 607545 Methylsterol monooxygenase 1 (EC (C-4 methylsterol oxidase) (Sterol-C4-methyl oxidase) [DESP4] [ERG25] [SC4MOL]
MSX1          4p16.1-p16.3    P28360     MSX1_HUMAN  142983 Homeobox protein MSX-1 (Homeobox protein Hox-7) (Msh homeobox 1-like protein) [HOX7]
MTHFD2L       4q13.3          Q9H903     MTD2L_HUMAN 614047 Probable bifunctional methylenetetrahydrofolate dehydrogenase/cyclohydrolase 2 (NADP-dependent methylenetetrahydrofolate dehydrogenase 2-like protein) (MTHFD2-like) [Includes: NAD-dependent methylenetetrahydrofolate dehydrogenase (EC; Methenyltetrahydrofolate cyclohydrolase (EC]
MTNR1A        4q35.1          P48039     MTR1A_HUMAN 600665 Melatonin receptor type 1A (Mel-1A-R) (Mel1a receptor)
MTRNR2L13     4q32.1          S4R3P1     HMN13_HUMAN        Humanin-like 13 (HN13) (MT-RNR2-like protein 13)
MTTP          4q24            P55157     MTP_HUMAN   157147 Microsomal triglyceride transfer protein large subunit precursor [MTP]
MUC7          4q13-q21        Q8TAX7     MUC7_HUMAN  158375 Mucin-7 precursor (MUC-7) (Apo-MG2) (Salivary mucin-7) [MG2]
MXD4          4p16.3          Q14582     MAD4_HUMAN         Max dimerization protein 4 (Max dimerizer 4) (Class C basic helix-loop-helix protein 12) (bHLHc12) (Max-associated protein 4) (Max-interacting transcriptional repressor MAD4) [BHLHC12] [MAD4]
MYL5          4p16.3          Q02045     MYL5_HUMAN  160782 Myosin light chain 5 (Myosin regulatory light chain 5) (Superfast myosin regulatory light chain 2) (MYLC2) (MyLC-2)
MYOZ2         4q26-q27        Q9NPC6     MYOZ2_HUMAN 605602 Myozenin-2 (Calsarcin-1) (FATZ-related protein 2) [C4orf5]
N4BP2         4p14            Q86UW6     N4BP2_HUMAN 619139 NEDD4-binding protein 2 (EC 3.-.-.-) (N4BP2) (BCL-3-binding protein) [B3BP] [KIAA1413]
NAA11         4q21.23         Q9BSU3     NAA11_HUMAN        N-alpha-acetyltransferase 11 (EC (N-terminal acetyltransferase complex ARD1 subunit homolog B) (hARD2) (NatA catalytic subunit Naa11) [ARD1B] [ARD2]
NAA15         4q31.1          Q9BXJ9     NAA15_HUMAN 608000 N-alpha-acetyltransferase 15, NatA auxiliary subunit (Gastric cancer antigen Ga19) (N-terminal acetyltransferase) (NMDA receptor-regulated protein 1) (Protein tubedown-1) (Tbdn100) [GA19] [NARG1] [NATH] [TBDN100]
NAAA          4q21.1          Q02083     NAAA_HUMAN  607469 N-acylethanolamine-hydrolyzing acid amidase precursor (EC (Acid ceramidase-like protein) (Acylsphingosine deacylase NAAA) (EC (N-acylsphingosine amidohydrolase-like) (ASAH-like protein) [Contains: N-acylethanolamine-hydrolyzing acid amidase subunit alpha; N-acylethanolamine-hydrolyzing acid amidase subunit beta] [ASAHL] [PLT]
NAF1          4q32.2          Q96HR8     NAF1_HUMAN  617868 H/ACA ribonucleoprotein complex non-core subunit NAF1 (hNAF1)
NAP1L5        4q21-q22        Q96NT1     NP1L5_HUMAN 612203 Nucleosome assembly protein 1-like 5 (Down-regulated in liver malignancy) [DRLM]
NAT8L         4p16.3          Q8N9F0     NAT8L_HUMAN 610647 N-acetylaspartate synthetase (EC (NAA synthetase) (Camello-like protein 3) (N-acetyltransferase 8-like protein) [CML3]
NCAPG         4p15.33         Q9BPX3     CND3_HUMAN  606280 Condensin complex subunit 3 (Chromosome-associated protein G) (Condensin subunit CAP-G) (hCAP-G) (Melanoma antigen NY-MEL-3) (Non-SMC condensin I complex subunit G) (XCAP-G homolog) [CAPG] [NYMEL3]
NDNF          4q27            Q8TB73     NDNF_HUMAN  616506 Protein NDNF precursor (Neuron-derived neurotrophic factor) [C4orf31] [UNQ2748/PRO6487]
NDST3         4q26            O95803     NDST3_HUMAN 603950 Bifunctional heparan sulfate N-deacetylase/N-sulfotransferase 3 (EC (Glucosaminyl N-deacetylase/N-sulfotransferase 3) (NDST-3) (hNDST-3) (N-heparan sulfate sulfotransferase 3) (N-HSST 3) [Includes: Heparan sulfate N-deacetylase 3 (EC 3.-.-.-); Heparan sulfate N-sulfotransferase 3 (EC 2.8.2.-)] [HSST3] [UNQ2544/PRO4998]
NDST4         4q25-q26        Q9H3R1     NDST4_HUMAN 615039 Bifunctional heparan sulfate N-deacetylase/N-sulfotransferase 4 (EC (Glucosaminyl N-deacetylase/N-sulfotransferase 4) (NDST-4) (N-heparan sulfate sulfotransferase 4) (N-HSST 4) [Includes: Heparan sulfate N-deacetylase 4 (EC 3.-.-.-); Heparan sulfate N-sulfotransferase 4 (EC 2.8.2.-)] [HSST4]
NDUFC1        4q28.2-q28.3    O43677     NDUC1_HUMAN 603844 NADH dehydrogenase [ubiquinone] 1 subunit C1, mitochondrial precursor (Complex I-KFYI) (CI-KFYI) (NADH-ubiquinone oxidoreductase KFYI subunit)
NEIL3         4q34.3          Q8TAT5     NEIL3_HUMAN 608934 Endonuclease 8-like 3 (EC 3.2.2.-) (EC (DNA glycosylase FPG2) (DNA glycosylase/AP lyase Neil3) (Endonuclease VIII-like 3) (Nei-like protein 3)
NEK1          4q33            Q96PY6     NEK1_HUMAN  604588 Serine/threonine-protein kinase Nek1 (EC (Never in mitosis A-related kinase 1) (NimA-related protein kinase 1) (Renal carcinoma antigen NY-REN-55) [KIAA1901]
NELFA         4p16.3          Q9H3P2     NELFA_HUMAN 606026 Negative elongation factor A (NELF-A) (Wolf-Hirschhorn syndrome candidate 2 protein) [WHSC2] [P/OKcl.15]
NEUROG2       4q25            Q9H2A3     NGN2_HUMAN  606624 Neurogenin-2 (NGN-2) (Class A basic helix-loop-helix protein 8) (bHLHa8) (Protein atonal homolog 4) [ATOH4] [BHLHA8] [NGN2]
NFKB1         4q24            P19838     NFKB1_HUMAN 164011 Nuclear factor NF-kappa-B p105 subunit (DNA-binding factor KBF1) (EBP-1) (Nuclear factor of kappa light polypeptide gene enhancer in B-cells 1) [Contains: Nuclear factor NF-kappa-B p50 subunit]
NFXL1         4p12            Q6ZNB6     NFXL1_HUMAN        NF-X1-type zinc finger protein NFXL1 (Ovarian zinc finger protein) (hOZFP) [OZFP]
NIPAL1        4p12            Q6NVV3     NIPA3_HUMAN        Magnesium transporter NIPA3 (NIPA-like protein 1) (Non-imprinted in Prader-Willi/Angelman syndrome region protein 3) [NIPA3] [NPAL1]
NKX1-1        4p16.3          Q15270     NKX11_HUMAN 617869 NK1 transcription factor-related protein 1 (Homeobox protein 153) (HPX-153) (Homeobox protein SAX-2) (NKX-1.1) [HPX153] [SAX2]
NKX3-2        4p16.1          P78367     NKX32_HUMAN 602183 Homeobox protein Nkx-3.2 (Bagpipe homeobox protein homolog 1) (Homeobox protein NK-3 homolog B) [BAPX1] [NKX3B]
NKX6-1        4q21.2-q22      P78426     NKX61_HUMAN 602563 Homeobox protein Nkx-6.1 (Homeobox protein NK-6 homolog A) [NKX6A]
NMU           4q11            P48645     NMU_HUMAN   605103 Neuromedin-U precursor [Contains: Neuromedin precursor-related peptide 36 (NURP36); Neuromedin precursor-related peptide 33 (NURP33); Neuromedin-U-25 (NmU-25)]
NOA1          4q12            Q8NC60     NOA1_HUMAN  614919 Nitric oxide-associated protein 1 [C4orf14]
NOCT          4q31.1          Q9UK39     NOCT_HUMAN  608468 Nocturnin precursor (EC (Carbon catabolite repression 4-like protein) [CCR4] [CCRN4L] [NOC]
NOP14         4p16.3          P78316     NOP14_HUMAN 611526 Nucleolar protein 14 (Nucleolar complex protein 14) [C4orf9] [NOL14] [RES4-25]
NPFFR2        4q21            Q9Y5X5     NPFF2_HUMAN 607449 Neuropeptide FF receptor 2 (G-protein coupled receptor 74) (G-protein coupled receptor HLWAR77) (Neuropeptide G-protein coupled receptor) [GPR74] [NPFF2] [NPGPR]
NPNT          4q25            Q6UXI9     NPNT_HUMAN  610306 Nephronectin precursor (Preosteoblast EGF-like repeat protein with MAM domain) (Protein EGFL6-like) [EGFL6L] [POEM] [UNQ295/PRO334]
NPY1R         4q31.3-q32      P25929     NPY1R_HUMAN 162641 Neuropeptide Y receptor type 1 (NPY1-R) [NPYR] [NPYY1]
NPY2R         4q31            P49146     NPY2R_HUMAN 162642 Neuropeptide Y receptor type 2 (NPY2-R) (NPY-Y2 receptor) (Y2 receptor)
NPY5R         4q31-q32        Q15761     NPY5R_HUMAN 602001 Neuropeptide Y receptor type 5 (NPY5-R) (NPY-Y5 receptor) (NPYY5-R) (Y5 receptor) [NPYR5]
NR3C2         4q31.1          P08235     MCR_HUMAN   600983 Mineralocorticoid receptor (MR) (Nuclear receptor subfamily 3 group C member 2) [MCR] [MLR]
NSD2          4p16.3          O96028     NSD2_HUMAN  602952 Histone-lysine N-methyltransferase NSD2 (EC (Multiple myeloma SET domain-containing protein) (MMSET) (Nuclear SET domain-containing protein 2) (Protein trithorax-5) (Wolf-Hirschhorn syndrome candidate 1 protein) [KIAA1090] [MMSET] [TRX5] [WHSC1]
NSG1          4p16.3          P42857     NSG1_HUMAN  607645 Neuronal vesicle trafficking-associated protein 1 (Neuron-enriched endosomal protein of 21 kDa) (Neuron-specific protein family member 1) [D4S234] [NEEP21]
NSUN7         4p14            Q8NE18     NSUN7_HUMAN        Putative methyltransferase NSUN7 (EC 2.1.1.-) (NOL1/NOP2/Sun domain family member 7)
NUDT6         4q26            P53370     NUDT6_HUMAN 606261 Nucleoside diphosphate-linked moiety X motif 6 (EC 3.6.1.-) (Nudix motif 6) (Antisense basic fibroblast growth factor) (Protein GFG) [FGF2AS]
NUDT9         4q22.1          Q9BW91     NUDT9_HUMAN 606022 ADP-ribose pyrophosphatase, mitochondrial precursor (EC (ADP-ribose diphosphatase) (ADP-ribose phosphohydrolase) (Adenosine diphosphoribose pyrophosphatase) (ADPR-PPase) (Nucleoside diphosphate-linked moiety X motif 9) (Nudix motif 9) [NUDT10] [PSEC0099] [UNQ3012/PRO9771]
NUP54         4q21.21         Q7Z3B4     NUP54_HUMAN 607607 Nucleoporin p54 (54 kDa nucleoporin)
NWD2          4p14            Q9ULI1     NWD2_HUMAN         NACHT and WD repeat domain-containing protein 2 (Leucine-rich repeat and WD repeat-containing protein KIAA1239) [KIAA1239]
OCIAD1        4p11            Q9NX40     OCAD1_HUMAN        OCIA domain-containing protein 1 (Ovarian cancer immunoreactive antigen domain containing 1) (Ovarian carcinoma immunoreactive antigen) [ASRIJ] [OCIA]
OCIAD2        4p12            Q56VL3     OCAD2_HUMAN        OCIA domain-containing protein 2 (Ovarian carcinoma immunoreactive antigen-like protein)
ODAM          4q13.3          A1E959     ODAM_HUMAN  614843 Odontogenic ameloblast-associated protein precursor (Apin) [APIN]
ODAPH         4q21.1          Q17RF5     ODAPH_HUMAN 614829 Odontogenesis associated phosphoprotein precursor [C4orf26]
OPRPN         4q13.2          Q99935     PROL1_HUMAN 608936 Opiorphin prepropeptide precursor (Basic proline-rich lacrimal protein) (Proline-rich protein 1) (PRL1) [Contains: Opiorphin] [BPLP] [PROL1]
OSTC          4q25            Q9NRP0     OSTC_HUMAN  619023 Oligosaccharyltransferase complex subunit OSTC (Hydrophobic protein HSF-28) [DC2] [HDCMD45P] [HSPC307]
OTOP1         4p16.2          Q7RTM1     OTOP1_HUMAN 607806 Proton channel OTOP1 (Otopetrin-1) (hOtop1)
OTUD4         4q31.21         Q01804     OTUD4_HUMAN 611744 OTU domain-containing protein 4 (EC (HIV-1-induced protein HIN-1) [HIN-1] [KIAA1046]
PABPC4L       4q28.3          P0CB38     PAB4L_HUMAN        Polyadenylate-binding protein 4-like (PABP-4-like) (Poly(A)-binding protein 4-like)
PACRGL        4p15.31         Q8N7B6     PACRL_HUMAN        PACRG-like protein [C4orf28]
PAICS         4pter-q21       P22234     PUR6_HUMAN  172439 Multifunctional protein ADE2 [Includes: Phosphoribosylaminoimidazole-succinocarboxamide synthase (EC (SAICAR synthetase); Phosphoribosylaminoimidazole carboxylase (EC (AIR carboxylase) (AIRC)] [ADE2] [AIRC] [PAIS]
PALLD         4q32.3          Q8WX93     PALLD_HUMAN 608092 Palladin (SIH002) (Sarcoma antigen NY-SAR-77) [KIAA0992] [CGI-151]
PAPSS1        4q24            O43252     PAPS1_HUMAN 603262 Bifunctional 3'-phosphoadenosine 5'-phosphosulfate synthase 1 (PAPS synthase 1) (PAPSS 1) (Sulfurylase kinase 1) (SK 1) (SK1) [Includes: Sulfate adenylyltransferase (EC (ATP-sulfurylase) (Sulfate adenylate transferase) (SAT); Adenylyl-sulfate kinase (EC (3'-phosphoadenosine-5'-phosphosulfate synthase) (APS kinase) (Adenosine-5'-phosphosulfate 3'-phosphotransferase) (Adenylylsulfate 3'-phosphotransferase)] [ATPSK1] [PAPSS]
PAQR3         4q21.21         Q6TCH7     PAQR3_HUMAN 614577 Progestin and adipoQ receptor family member 3 (Progestin and adipoQ receptor family member III) (Raf kinase trapping to Golgi) (RKTG)
PARM1         4q13.3          Q6UWI2     PARM1_HUMAN 617688 Prostate androgen-regulated mucin-like protein 1 precursor (PARM-1) [UNQ1879/PRO4322]
PCDH10        4q28.3          Q9P2E7     PCD10_HUMAN 608286 Protocadherin-10 precursor [KIAA1400]
PCDH18        4q31            Q9HCL0     PCD18_HUMAN 608287 Protocadherin-18 precursor [KIAA1562]
PCDH7         4p15            O60245     PCDH7_HUMAN 602988 Protocadherin-7 precursor (Brain-heart protocadherin) (BH-Pcdh) [BHPCDH]
PCGF3         4p16.3          Q3KNV8     PCGF3_HUMAN        Polycomb group RING finger protein 3 (RING finger protein 3A) [RNF3] [RNF3A]
PDCL2         4q12            Q8N4E4     PDCL2_HUMAN 611676 Phosducin-like protein 2
PDE5A         4q25-q27        O76074     PDE5A_HUMAN 603310 cGMP-specific 3',5'-cyclic phosphodiesterase (EC (cGMP-binding cGMP-specific phosphodiesterase) (CGB-PDE) [PDE5]
PDE6B         4p16.3          P35913     PDE6B_HUMAN 180072 Rod cGMP-specific 3',5'-cyclic phosphodiesterase subunit beta precursor (EC (GMP-PDE beta) [PDEB]
PDGFC         4q32            Q9NRA1     PDGFC_HUMAN 608452 Platelet-derived growth factor C precursor (PDGF-C) (Fallotein) (Spinal cord-derived growth factor) (SCDGF) (VEGF-E) [Contains: Platelet-derived growth factor C, latent form (PDGFC latent form); Platelet-derived growth factor C, receptor-binding form (PDGFC receptor-binding form)] [SCDGF] [UNQ174/PRO200]
PDGFRA        4q11-q13        P16234     PGFRA_HUMAN 173490 Platelet-derived growth factor receptor alpha precursor (EC (PDGF-R-alpha) (PDGFR-alpha) (Alpha platelet-derived growth factor receptor) (Alpha-type platelet-derived growth factor receptor) (CD140 antigen-like family member A) (CD140a antigen) (Platelet-derived growth factor alpha receptor) (Platelet-derived growth factor receptor 2) (PDGFR-2) (CD140a antigen) [PDGFR2] [RHEPDGFRA]
PDHA2         4q22-q23        P29803     ODPAT_HUMAN 179061 Pyruvate dehydrogenase E1 component subunit alpha, testis-specific form, mitochondrial precursor (EC (PDHE1-A type II) [PDHAL]
PDLIM3        4q35            Q53GG5     PDLI3_HUMAN 605889 PDZ and LIM domain protein 3 (Actinin-associated LIM protein) (Alpha-actinin-2-associated LIM protein) [ALP]
PDLIM5        4q22            Q96HC4     PDLI5_HUMAN 605904 PDZ and LIM domain protein 5 (Enigma homolog) (Enigma-like PDZ and LIM domains protein) [ENH] [L9]
PDS5A         4p14            Q29RF7     PDS5A_HUMAN 613200 Sister chromatid cohesion protein PDS5 homolog A (Cell proliferation-inducing gene 54 protein) (Sister chromatid cohesion protein 112) (SCC-112) [KIAA0648] [PDS5] [PIG54]
PF4           4q12-q13        P02776     PLF4_HUMAN  173460 Platelet factor 4 precursor (PF-4) (C-X-C motif chemokine 4) (Iroplact) (Oncostatin-A) [Contains: Platelet factor 4, short form (Endothelial cell growth inhibitor)] [CXCL4] [SCYB4]
PF4V1         4q12-q21        P10720     PF4V_HUMAN  173461 Platelet factor 4 variant precursor (C-X-C motif chemokine 4 variant) (CXCL4L1) (PF4alt) (PF4var1) [Contains: Platelet factor 4 variant(4-74); Platelet factor 4 variant(5-74); Platelet factor 4 variant(6-74)] [CXCL4V1] [SCYB4V1]
PGM2          4p14-q12        Q96G03     PGM2_HUMAN  172000 Phosphoglucomutase-2 (EC (PGM 2) (Glucose phosphomutase 2) (Phosphodeoxyribomutase) (Phosphopentomutase) (EC [MSTP006]
PGRMC2        4q26            O15173     PGRC2_HUMAN 607735 Membrane-associated progesterone receptor component 2 (Progesterone membrane-binding protein) (Steroid receptor protein DG6) [DG6] [PMBP]
PHOX2B        4p12            Q99453     PHX2B_HUMAN 603851 Paired mesoderm homeobox protein 2B (Neuroblastoma Phox) (NBPhox) (PHOX2B homeodomain protein) (Paired-like homeobox 2B) [PMX2B]
PI4K2B        4p15.31         Q8TCG2     P4K2B_HUMAN 612101 Phosphatidylinositol 4-kinase type 2-beta (EC (Phosphatidylinositol 4-kinase type II-beta) (PI4KII-BETA)
PIGG          4p16.3          Q5H8A4     PIGG_HUMAN  616918 GPI ethanolamine phosphate transferase 2 (EC 2.-.-.-) (GPI7 homolog) (hGPI7) (Phosphatidylinositol-glycan biosynthesis class G protein) (PIG-G) [GPI7] [UNQ1930/PRO4405]
PIGY          4q22.1          Q3MUY2     PIGY_HUMAN  610662 Phosphatidylinositol N-acetylglucosaminyltransferase subunit Y (Phosphatidylinositol-glycan biosynthesis class Y protein) (PIG-Y)
PITX2         4q25-q27        Q99697     PITX2_HUMAN 601542 Pituitary homeobox 2 (ALL1-responsive protein ARP1) (Homeobox protein PITX2) (Paired-like homeodomain transcription factor 2) (RIEG bicoid-related homeobox transcription factor) (Solurshin) [ARP1] [RGS] [RIEG] [RIEG1]
PKD2          4q21-q23        Q13563     PKD2_HUMAN  173910 Polycystin-2 (PC2) (Autosomal dominant polycystic kidney disease type II protein) (Polycystic kidney disease 2 protein) (Polycystwin) (R48321) (Transient receptor potential cation channel subfamily P member 2) [TRPP2]
PLA2G12A      4q25            Q9BZM1     PG12A_HUMAN 611652 Group XIIA secretory phospholipase A2 precursor (EC (GXII sPLA2) (sPLA2-XII) (Phosphatidylcholine 2-acylhydrolase 12A) [PLA2G12] [FKSG38] [UNQ2519/PRO6012]
PLAC8         4q21.3          Q9NZF1     PLAC8_HUMAN 607515 Placenta-specific gene 8 protein (Protein C15) [BM-004]
PLK4          4q28.1          O00444     PLK4_HUMAN  605031 Serine/threonine-protein kinase PLK4 (EC (Polo-like kinase 4) (PLK-4) (Serine/threonine-protein kinase 18) (Serine/threonine-protein kinase Sak) [SAK] [STK18]
PLRG1         4q31.3          O43660     PLRG1_HUMAN 605961 Pleiotropic regulator 1
POLN          4p16.3          Q7Z5Q5     DPOLN_HUMAN 610887 DNA polymerase nu (EC
POLR2B        4q12            P30876     RPB2_HUMAN  180661 DNA-directed RNA polymerase II subunit RPB2 (EC (DNA-directed RNA polymerase II 140 kDa polypeptide) (DNA-directed RNA polymerase II subunit B) (RNA polymerase II subunit 2) (RNA polymerase II subunit B2)
POU4F2        4q31.2          Q12837     PO4F2_HUMAN 113725 POU domain, class 4, transcription factor 2 (Brain-specific homeobox/POU domain protein 3B) (Brain-3B) (Brn-3B) [BRN3B]
PPA2          4q24            Q9H2U2     IPYR2_HUMAN 609988 Inorganic pyrophosphatase 2, mitochondrial precursor (EC (Pyrophosphatase SID6-306) (Pyrophosphate phospho-hydrolase 2) (PPase 2) [HSPC124]
PPARGC1A      4p15.1          Q9UBK2     PRGC1_HUMAN 604517 Peroxisome proliferator-activated receptor gamma coactivator 1-alpha (PGC-1-alpha) (PPAR-gamma coactivator 1-alpha) (PPARGC-1-alpha) (Ligand effect modulator 6) [LEM6] [PGC1] [PGC1A] [PPARGC1]
PPAT          4q12            Q06203     PUR1_HUMAN  172450 Amidophosphoribosyltransferase precursor (EC (ATase) (Glutamine phosphoribosylpyrophosphate amidotransferase) (GPAT) [GPAT]
PPBP          4q12-q13        P02775     CXCL7_HUMAN 121010 Platelet basic protein precursor (PBP) (C-X-C motif chemokine 7) (Leukocyte-derived growth factor) (LDGF) (Macrophage-derived growth factor) (MDGF) (Small-inducible cytokine B7) [Contains: Connective tissue-activating peptide III (CTAP-III) (LA-PF4) (Low-affinity platelet factor IV); TC-2; Connective tissue-activating peptide III(1-81) (CTAP-III(1-81)); Beta-thromboglobulin (Beta-TG); Neutrophil-activating peptide 2(74) (NAP-2(74)); Neutrophil-activating peptide 2(73) (NAP-2(73)); Neutrophil-activating peptide 2 (NAP-2); TC-1; Neutrophil-activating peptide 2(1-66) (NAP-2(1-66)); Neutrophil-activating peptide 2(1-63) (NAP-2(1-63))] [CTAP3] [CXCL7] [SCYB7] [TGB1] [THBGB1]
PPEF2         4q21.21         O14830     PPE2_HUMAN  602256 Serine/threonine-protein phosphatase with EF-hands 2 (EC (PPEF-2)
PPID          4q31.3          Q08752     PPID_HUMAN  601753 Peptidyl-prolyl cis-trans isomerase D (EC (PPIase D) (40 kDa peptidyl-prolyl cis-trans isomerase) (Cyclophilin-40) (CYP-40) (Cyclophilin-related protein) (Rotamase D) [CYP40] [CYPD]
PPM1K         4q22.1          Q8N3J5     PPM1K_HUMAN 611065 Protein phosphatase 1K, mitochondrial precursor (EC (PP2C domain-containing protein phosphatase 1K) (PP2C-like mitochondrial protein) (PP2C-type mitochondrial phosphoprotein phosphatase) (PTMP) (Protein phosphatase 2C isoform kappa) (PP2C-kappa) [PP2CM]
PPP2R2C       4p16            Q9Y2T4     2ABG_HUMAN  605997 Serine/threonine-protein phosphatase 2A 55 kDa regulatory subunit B gamma isoform (IMYPNO1) (PP2A subunit B isoform B55-gamma) (PP2A subunit B isoform PR55-gamma) (PP2A subunit B isoform R2-gamma) (PP2A subunit B isoform gamma)
PPP3CA        4q21-q24        Q08209     PP2BA_HUMAN 114105 Serine/threonine-protein phosphatase 2B catalytic subunit alpha isoform (EC (CAM-PRP catalytic subunit) (Calmodulin-dependent calcineurin A subunit alpha isoform) [CALNA] [CNA]
PRDM5         4q25-q26        Q9NQX1     PRDM5_HUMAN 614161 PR domain zinc finger protein 5 (EC 2.1.1.-) (PR domain-containing protein 5) [PFM2]
PRDM8         4q21            Q9NQV8     PRDM8_HUMAN 616639 PR domain zinc finger protein 8 (EC 2.1.1.-) (PR domain-containing protein 8) [PFM5]
PRIMPOL       4q35.1          Q96LW4     PRIPO_HUMAN 615421 DNA-directed primase/polymerase protein (EC 2.7.7.-) (hPrimpol1) (Coiled-coil domain-containing protein 111) [CCDC111]
PRKG2         4q13.1-q21.1    Q13237     KGP2_HUMAN  601591 cGMP-dependent protein kinase 2 (EC (cGK 2) (cGK2) (cGMP-dependent protein kinase II) (cGKII) [PRKGR2]
PRMT9         4q31.23         Q6P2P2     ANM9_HUMAN  616125 Protein arginine N-methyltransferase 9 (Protein arginine N-methyltransferase 10) (EC [PRMT10]
PROM1         4p15.33         O43490     PROM1_HUMAN 604365 Prominin-1 precursor (Antigen AC133) (Prominin-like protein 1) (CD133 antigen) [PROML1] [MSTP061]
PRR27         4q13.3          Q6MZM9     PRR27_HUMAN        Proline-rich protein 27 precursor [C4orf40]
PRSS12        4q28.1          P56730     NETR_HUMAN  606709 Neurotrypsin precursor (EC 3.4.21.-) (Leydin) (Motopsin) (Serine protease 12)
PRSS48        4q31.3          Q7RTY5     PRS48_HUMAN        Serine protease 48 precursor (EC 3.4.21.-) (Epidermis-specific serine protease-like protein) [ESSPL]
PSAPL1        4p16.1          Q6NUJ1     SAPL1_HUMAN        Proactivator polypeptide-like 1 precursor [Contains: Saposin A-like; Saposin B-Val-like; Saposin B-like; Saposin C-like; Saposin D-like]
PTPN13        4q21.3          Q12923     PTN13_HUMAN 600267 Tyrosine-protein phosphatase non-receptor type 13 (EC (Fas-associated protein-tyrosine phosphatase 1) (FAP-1) (PTP-BAS) (Protein-tyrosine phosphatase 1E) (PTP-E1) (hPTPE1) (Protein-tyrosine phosphatase PTPL1) [PNP1] [PTP1E] [PTPL1]
PTTG2         4p12            Q9NZH5     PTTG2_HUMAN 604231 Securin-2 (Pituitary tumor-transforming gene 2 protein)
PYURF         4q22.1          Q96I23     PREY_HUMAN         Protein preY, mitochondrial precursor (PIGY upstream reading frame protein) [PREY]
QDPR          4p15.31         P09417     DHPR_HUMAN  612676 Dihydropteridine reductase (EC (HDHPR) (Quinoid dihydropteridine reductase) (Short chain dehydrogenase/reductase family 33C member 1) [DHPR] [SDR33C1]
QRFPR         4q27            Q96P65     QRFPR_HUMAN 606925 Pyroglutamylated RF-amide peptide receptor (AQ27) (G-protein coupled receptor 103) (Orexigenic neuropeptide QRFP receptor) (SP9155) [GPR103]
RAB28         4p16.1          P51157     RAB28_HUMAN 612994 Ras-related protein Rab-28 precursor
RAB33B        4q28            Q9H082     RB33B_HUMAN 605950 Ras-related protein Rab-33B
RAP1GDS1      4q21            P52306     GDS1_HUMAN  179502 Rap1 GTPase-GDP dissociation stimulator 1 (Exchange factor smgGDS) (SMG GDS protein) (SMG P21 stimulatory GDP/GTP exchange protein)
RAPGEF2       4q32.1          Q9Y4G8     RPGF2_HUMAN 609530 Rap guanine nucleotide exchange factor 2 (Cyclic nucleotide ras GEF) (CNrasGEF) (Neural RAP guanine nucleotide exchange protein) (nRap GEP) (PDZ domain-containing guanine nucleotide exchange factor 1) (PDZ-GEF1) (RA-GEF-1) (Ras/Rap1-associating GEF-1) [KIAA0313] [NRAPGEP] [PDZGEF1]
RASGEF1B      4q21.21         Q0VAM2     RGF1B_HUMAN 614532 Ras-GEF domain-containing family member 1B (GPI gamma-4) [GPIG4]
RASL11B       4q12            Q9BPW5     RSLBB_HUMAN 612404 Ras-like protein family member 11B (EC
RASSF6        4q21.1          Q6ZTQ3     RASF6_HUMAN 612620 Ras association domain-containing protein 6
RBM46         4q32.1          Q8TBY0     RBM46_HUMAN        Probable RNA-binding protein 46 (Cancer/testis antigen 68) (CT68) (RNA-binding motif protein 46)
RBM47         4p14            A0AV96     RBM47_HUMAN 619104 RNA-binding protein 47 (RNA-binding motif protein 47)
RBPJ          4p15.2          Q06330     SUH_HUMAN   147183 Recombining binding protein suppressor of hairless (CBF-1) (J kappa-recombination signal-binding protein) (RBP-J kappa) (RBP-J) (RBP-JK) (Renal carcinoma antigen NY-REN-30) [IGKJRB] [IGKJRB1] [RBPJK] [RBPSUH]
RCHY1         4q21.1          Q96PM5     ZN363_HUMAN 607680 RING finger and CHY zinc finger domain-containing protein 1 (EC (Androgen receptor N-terminal-interacting protein) (CH-rich-interacting match with PLAG1) (E3 ubiquitin-protein ligase Pirh2) (RING finger protein 199) (RING-type E3 ubiquitin transferase RCHY1) (Zinc finger protein 363) (p53-induced RING-H2 protein) (hPirh2) [ARNIP] [CHIMP] [PIRH2] [RNF199] [ZNF363]
REELD1        4q31.22         A0A1B0GV85 RELD1_HUMAN        Reelin domain-containing protein 1 precursor
RELL1         4p14            Q8IUW5     RELL1_HUMAN 611212 RELT-like protein 1 precursor [PSEC0162]
REST          4q12            Q13127     REST_HUMAN  600571 RE1-silencing transcription factor (Neural-restrictive silencer factor) (X2 box repressor) [NRSF] [XBR]
RFC1          4p13-p14        P35251     RFC1_HUMAN  102579 Replication factor C subunit 1 (Activator 1 140 kDa subunit) (A1 140 kDa subunit) (Activator 1 large subunit) (Activator 1 subunit 1) (DNA-binding protein PO-GA) (Replication factor C 140 kDa subunit) (RF-C 140 kDa subunit) (RFC140) (Replication factor C large subunit) [RFC140]
RGS12         4p16.3          O14924     RGS12_HUMAN 602512 Regulator of G-protein signaling 12 (RGS12)
RHOH          4p13            Q15669     RHOH_HUMAN  602037 Rho-related GTP-binding protein RhoH precursor (GTP-binding protein TTF) (Translocation three four protein) [ARHH] [TTF]
RNF150        4q31.21         Q9ULK6     RN150_HUMAN        RING finger protein 150 precursor [KIAA1214]
RNF175        4q31.3          Q8N4F7     RN175_HUMAN        RING finger protein 175
RNF212        4p16.3          Q495C1     RN212_HUMAN 612041 Probable E3 SUMO-protein ligase RNF212 (EC 2.3.2.-) (Probable E3 SUMO-protein transferase RNF212) (RING finger protein 212)
RNF4          4p16.3          P78317     RNF4_HUMAN  602850 E3 ubiquitin-protein ligase RNF4 (EC (RING finger protein 4) (RING-type E3 ubiquitin transferase RNF4) (Small nuclear ring finger protein) (Protein SNURF) [SNURF] [RES4-26]
RPL34         4q24            P49207     RL34_HUMAN  616862 60S ribosomal protein L34 (Large ribosomal subunit protein eL34)
RPL9          4p13            P32969     RL9_HUMAN   603686 60S ribosomal protein L9 (Large ribosomal subunit protein uL6) [OK/SW-cl.103]
RPS3A         4q31.2-q31.3    P61247     RS3A_HUMAN  180478 40S ribosomal protein S3a (Small ribosomal subunit protein eS1) (v-fos transformation effector protein) (Fte-1) [FTE1] [MFTL]
RRH           4q25            O14718     OPSX_HUMAN  605224 Visual pigment-like receptor peropsin
RUFY3         4q13.3          Q7L099     RUFY3_HUMAN 611194 Protein RUFY3 (RUN and FYVE domain-containing protein 3) (Rap2-interacting protein x) (RIPx) (Single axon-regulated protein) (Singar) [KIAA0871]
RWDD4         4q35.1          Q6NW29     RWDD4_HUMAN        RWD domain-containing protein 4 (Protein FAM28A) [FAM28A] [RWDD4A]
RXFP1         4q31.3          Q9HBX9     RXFP1_HUMAN 606654 Relaxin receptor 1 (Leucine-rich repeat-containing G-protein coupled receptor 7) (Relaxin family peptide receptor 1) [LGR7]
S100P         4p16            P25815     S100P_HUMAN 600614 Protein S100-P (Migration-inducing gene 9 protein) (MIG9) (Protein S100-E) (S100 calcium-binding protein P) [S100E]
SAP30         4q34.1          O75446     SAP30_HUMAN 603378 Histone deacetylase complex subunit SAP30 (30 kDa Sin3-associated polypeptide) (Sin3 corepressor complex subunit SAP30) (Sin3-associated polypeptide p30)
SCARB2        4q21.1          Q14108     SCRB2_HUMAN 602257 Lysosome membrane protein 2 (85 kDa lysosomal membrane sialoglycoprotein) (LGP85) (CD36 antigen-like 2) (Lysosome membrane protein II) (LIMP II) (Scavenger receptor class B member 2) [CD36L2] [LIMP2] [LIMPII]
SCD5          4q21.22         Q86SK9     SCD5_HUMAN  608370 Stearoyl-CoA desaturase 5 (EC (Acyl-CoA-desaturase 4) (HSCD5) (Stearoyl-CoA 9-desaturase) (Stearoyl-CoA desaturase 2) [ACOD4] [SCD2] [SCD4]
SCFD2         4q12            Q8WU76     SCFD2_HUMAN        Sec1 family domain-containing protein 2 (Syntaxin-binding protein 1-like 1) [STXBP1L1]
SCLT1         4q28.2          Q96NL6     SCLT1_HUMAN 611399 Sodium channel and clathrin linker 1 (Sodium channel-associated protein 1) [SAP1]
SCOC          4q31.1          Q9UIL1     SCOC_HUMAN         Short coiled-coil protein [SCOCO] [HRIHFB2072]
SCRG1         4q31-q32        O75711     SCRG1_HUMAN 603163 Scrapie-responsive protein 1 precursor (Scrapie-responsive gene 1 protein) (ScRG-1) [UNQ390/PRO725]
SDAD1         4q21.1          Q9NVU7     SDA1_HUMAN         Protein SDA1 homolog (Nucleolar protein 130) (SDA1 domain-containing protein 1) (hSDA) [NUC130]
SEC24B        4q24            O95487     SC24B_HUMAN 607184 Protein transport protein Sec24B (SEC24-related protein B)
SEC24D        4q26            O94855     SC24D_HUMAN 607186 Protein transport protein Sec24D (SEC24-related protein D) [KIAA0755]
SEC31A        4q21.3          O94979     SC31A_HUMAN 610257 Protein transport protein Sec31A (ABP125) (ABP130) (SEC31-like protein 1) (SEC31-related protein A) (Web1-like protein) [KIAA0905] [SEC31L1] [HSPC275] [HSPC334]
SEL1L3        4p15.2          Q68CR1     SE1L3_HUMAN        Protein sel-1 homolog 3 (Suppressor of lin-12-like protein 3) (Sel-1L3) [KIAA0746]
SEPSECS       4p15.2          Q9HD40     SPCS_HUMAN  613009 O-phosphoseryl-tRNA(Sec) selenium transferase (EC (Liver-pancreas antigen) (LP) (SLA-p35) (SLA/LP autoantigen) (Selenocysteine synthase) (Sec synthase) (Selenocysteinyl-tRNA(Sec) synthase) (Sep-tRNA:Sec-tRNA synthase) (SepSecS) (Soluble liver antigen) (SLA) (UGA suppressor tRNA-associated protein) (tRNA(Ser/Sec)-associated antigenic protein) [TRNP48]
SEPTIN11      4q21            Q9NVA2     SEP11_HUMAN 612887 Septin-11 [SEPT11]
SETD7         4q28            Q8WTS6     SETD7_HUMAN 606594 Histone-lysine N-methyltransferase SETD7 (EC (Histone H3-K4 methyltransferase SETD7) (H3-K4-HMTase SETD7) (Lysine N-methyltransferase 7) (SET domain-containing protein 7) (SET7/9) [KIAA1717] [KMT7] [SET7] [SET9]
SFRP2         4q31.3          Q96HF1     SFRP2_HUMAN 604157 Secreted frizzled-related protein 2 precursor (FRP-2) (sFRP-2) (Secreted apoptosis-related protein 1) (SARP-1) [FRP2] [SARP1] [FKSG12] [UNQ361/PRO697]
SGCB          4q12            Q16585     SGCB_HUMAN  600900 Beta-sarcoglycan (Beta-SG) (43 kDa dystrophin-associated glycoprotein) (43DAG) (A3b)
SGMS2         4q25            Q8NHU3     SMS2_HUMAN  611574 Phosphatidylcholine:ceramide cholinephosphotransferase 2 (EC (Sphingomyelin synthase 2) [SMS2]
SH3BP2        4p16.3          P78314     3BP2_HUMAN  602104 SH3 domain-binding protein 2 (3BP-2) [3BP2] [RES4-23]
SH3D19        4q31.3          Q5HYK7     SH319_HUMAN 608674 SH3 domain-containing protein 19 (ADAM-binding protein Eve-1) (EEN-binding protein) (EBP)
SH3RF1        4q32.3          Q7Z6J0     SH3R1_HUMAN 618642 E3 ubiquitin-protein ligase SH3RF1 (EC (Plenty of SH3s) (Protein POSH) (RING finger protein 142) (RING-type E3 ubiquitin transferase SH3RF1) (SH3 domain-containing RING finger protein 1) (SH3 multiple domains protein 2) [KIAA1494] [POSH] [POSH1] [RNF142] [SH3MD2]
SH3TC1        4p16.1          Q8TE82     S3TC1_HUMAN        SH3 domain and tetratricopeptide repeat-containing protein 1
SHISA3        4p13            A0PJX4     SHSA3_HUMAN 617325 Protein shisa-3 homolog precursor
SHROOM3       4q21.1          Q8TF72     SHRM3_HUMAN 604570 Protein Shroom3 (Shroom-related protein) (hShrmL) [KIAA1481] [SHRML] [MSTP013]
SLAIN2        4p12            Q9P270     SLAI2_HUMAN 610492 SLAIN motif-containing protein 2 [KIAA1458]
SLBP          4p16.3          Q14493     SLBP_HUMAN  602422 Histone RNA hairpin-binding protein (Histone stem-loop-binding protein) [HBP]
SLC10A4       4p12            Q96EP9     NTCP4_HUMAN 618563 Sodium/bile acid cotransporter 4 (Na(+)/bile acid cotransporter 4) (Solute carrier family 10 member 4)
SLC10A6       4q22.1          Q3KNW5     SOAT_HUMAN  613366 Solute carrier family 10 member 6 (Sodium-dependent organic anion transporter) [SOAT]
SLC10A7       4q31.22         Q0GE19     NTCP7_HUMAN 611459 Sodium/bile acid cotransporter 7 (Na(+)/bile acid cotransporter 7) (Solute carrier family 10 member 7) [C4orf13] [P7] [PSEC0051]
SLC25A31      4q28            Q9H0C2     ADT4_HUMAN  610796 ADP/ATP translocase 4 (ADP,ATP carrier protein 4) (Adenine nucleotide translocator 4) (ANT 4) (Solute carrier family 25 member 31) (Sperm flagellar energy carrier protein) [AAC4] [ANT4] [SFEC]
SLC25A4       4q35            P12235     ADT1_HUMAN  103220 ADP/ATP translocase 1 (ADP,ATP carrier protein 1) (ADP,ATP carrier protein, heart/skeletal muscle isoform T1) (Adenine nucleotide translocator 1) (ANT 1) (Solute carrier family 25 member 4) [AAC1] [ANT1]
SLC26A1       4p16.3          Q9H2B4     S26A1_HUMAN 610130 Sulfate anion transporter 1 (SAT-1) (Solute carrier family 26 member 1) [SAT1]
SLC2A9        4p15.3-p16      Q9NRM0     GTR9_HUMAN  606142 Solute carrier family 2, facilitated glucose transporter member 9 (Glucose transporter type 9) (GLUT-9) (Urate transporter) [GLUT9]
SLC30A9       4p12-p13        Q6PML9     ZNT9_HUMAN  604604 Zinc transporter 9 (ZnT-9) (Human embryonic lung protein) (HuEL) (Solute carrier family 30 member 9) [C4orf1] [HUEL]
SLC34A2       4p15.2          O95436     NPT2B_HUMAN 604217 Sodium-dependent phosphate transport protein 2B (Sodium-phosphate transport protein 2B) (Na(+)-dependent phosphate cotransporter 2B) (NaPi3b) (Sodium/phosphate cotransporter 2B) (Na(+)/Pi cotransporter 2B) (NaPi-2b) (Solute carrier family 34 member 2)
SLC39A8       4q22-q24        Q9C0K1     S39A8_HUMAN 608732 Metal cation symporter ZIP8 precursor (BCG-induced integral membrane protein in monocyte clone 103 protein) (LIV-1 subfamily of ZIP zinc transporter 6) (LZT-Hs6) (Solute carrier family 39 member 8) (Zrt- and Irt-like protein 8) (ZIP-8) [BIGM103] [ZIP8] [PP3105]
SLC49A3       4p16.3          Q6UXD7     S49A3_HUMAN        Solute carrier family 49 member A3 (Major facilitator superfamily domain-containing protein 7) (Myosin light polypeptide 5 regulatory protein) (MYL5) [MFSD7] [LP2561] [UNQ385/PRO717]
SLC4A4        4q21            Q9Y6R1     S4A4_HUMAN  603345 Electrogenic sodium bicarbonate cotransporter 1 (Sodium bicarbonate cotransporter) (Na(+)/HCO3(-) cotransporter) (Solute carrier family 4 member 4) (kNBC1) [NBC] [NBC1] [NBCE1]
SLC7A11       4q28-q31        Q9UPY5     XCT_HUMAN   607933 Cystine/glutamate transporter (Amino acid transport system xc-) (Calcium channel blocker resistance protein CCBR1) (Solute carrier family 7 member 11) (xCT)
SLC9B1        4q24            Q4ZJI4     SL9B1_HUMAN 611527 Sodium/hydrogen exchanger 9B1 (Na(+)/H(+) exchanger-like domain-containing protein 1) (NHE domain-containing protein 1) (Sodium/hydrogen exchanger-like domain-containing protein 1) (Solute carrier family 9, subfamily B member 1) [NHA1] [NHEDC1]
SLC9B2        4q24            Q86UD5     SL9B2_HUMAN 611789 Sodium/hydrogen exchanger 9B2 (Na(+)/H(+) exchanger NHA2) (Na(+)/H(+) exchanger-like domain-containing protein 2) (NHE domain-containing protein 2) (Sodium/hydrogen exchanger-like domain-containing protein 2) (Solute carrier family 9 subfamily B member 2) [NHA2] [NHEDC2]
SLIT2         4p15.2          O94813     SLIT2_HUMAN 603746 Slit homolog 2 protein precursor (Slit-2) [Contains: Slit homolog 2 protein N-product; Slit homolog 2 protein C-product] [SLIL3]
SMAD1         4q31            Q15797     SMAD1_HUMAN 601595 Mothers against decapentaplegic homolog 1 (MAD homolog 1) (Mothers against DPP homolog 1) (JV4-1) (Mad-related protein 1) (SMAD family member 1) (SMAD 1) (Smad1) (hSMAD1) (Transforming growth factor-beta-signaling protein 1) (BSP-1) [BSP1] [MADH1] [MADR1]
SMARCA5       4q31.21         O60264     SMCA5_HUMAN 603375 SWI/SNF-related matrix-associated actin-dependent regulator of chromatin subfamily A member 5 (EC 3.6.4.-) (SWI/SNF-related matrix-associated actin-dependent regulator of chromatin A5) (Sucrose nonfermenting protein 2 homolog) (hSNF2H) [SNF2H] [WCRF135]
SMARCAD1      4q22-q23        Q9H4L7     SMRCD_HUMAN 612761 SWI/SNF-related matrix-associated actin-dependent regulator of chromatin subfamily A containing DEAD/H box 1 (EC (ATP-dependent helicase 1) (hHEL1) [KIAA1122]
SMIM14        4p14            Q96QK8     SIM14_HUMAN        Small integral membrane protein 14 [C4orf34]
SMIM20        4p15.2          Q8N5G0     SIM20_HUMAN 617465 Small integral membrane protein 20 (Mitochondrial translation regulation assembly intermediate of cytochrome c oxidase protein of 7 kDa) (MITRAC7) [Contains: Phoenixin-14 (PNX-14); Phoenixin-20 (PNX-20)] [C4orf52] [MITRAC7]
SMIM31        4q32.3          A0A1B0GVY4 SIM31_HUMAN        Small integral membrane protein 31
SMIM43        4q27            Q4W5P6     SIM43_HUMAN        Small integral membrane protein 43 [TMEM155]
SMR3A         4q13.3          Q99954     SMR3A_HUMAN 618340 Submaxillary gland androgen-regulated protein 3A precursor (Proline-rich protein 5) (Proline-rich protein PBI) [PBI] [PROL5]
SMR3B         4q13.3          P02814     SMR3B_HUMAN 611593 Submaxillary gland androgen-regulated protein 3B precursor (Proline-rich peptide P-B) (Proline-rich protein 3) [Contains: Peptide P-A; Peptide D1A] [PBII] [PRL3] [PROL3]
SNCA          4q21            P37840     SYUA_HUMAN  163890 Alpha-synuclein (Non-A beta component of AD amyloid) (Non-A4 component of amyloid precursor) (NACP) [NACP] [PARK1]
SNX25         4q35.1          Q9H3E2     SNX25_HUMAN        Sorting nexin-25 [MSTP043]
SOD3          4pter-q21       P08294     SODE_HUMAN  185490 Extracellular superoxide dismutase [Cu-Zn] precursor (EC (EC-SOD)
SORBS2        4q35.1          O94875     SRBS2_HUMAN 616349 Sorbin and SH3 domain-containing protein 2 (Arg-binding protein 2) (ArgBP2) (Arg/Abl-interacting protein 2) (Sorbin) [ARGBP2] [KIAA0777]
SORCS2        4p16.1          Q96PQ0     SORC2_HUMAN 606284 VPS10 domain-containing receptor SorCS2 precursor [Contains: SorCS2 122 kDa chain; SorCS2 104 kDa chain; SorCS2 18 kDa chain] [KIAA1329]
SOWAHB        4q21.1          A6NEL2     SWAHB_HUMAN        Ankyrin repeat domain-containing protein SOWAHB (Ankyrin repeat domain-containing protein 56) (Protein sosondowah homolog B) [ANKRD56]
SPARCL1       4q21.3          Q14515     SPRL1_HUMAN 606041 SPARC-like protein 1 precursor (High endothelial venule protein) (Hevin) (MAST 9)
SPATA18       4q11            Q8TC71     MIEAP_HUMAN 612814 Mitochondria-eating protein (Spermatogenesis-associated protein 18) [MIEAP]
SPATA4        4q34.2          Q8NEY3     SPAT4_HUMAN 609879 Spermatogenesis-associated protein 4 (Testis and spermatogenesis cell-related protein 2) (Testis spermatocyte apoptosis-related gene 2 protein) [TSARG2]
SPATA5        4q28.1          Q8NB90     AFG2H_HUMAN 613940 ATPase family protein 2 homolog (EC (Spermatogenesis-associated factor protein) (Spermatogenesis-associated protein 5) [AFG2] [SPAF]
SPCS3         4q34.3          P61009     SPCS3_HUMAN 618854 Signal peptidase complex subunit 3 (EC 3.4.-.-) (Microsomal signal peptidase 22/23 kDa subunit) (SPC22/23) (SPase 22/23 kDa subunit) [SPC22] [UNQ1841/PRO3567]
SPINK2        4q11            P20155     ISK2_HUMAN  605753 Serine protease inhibitor Kazal-type 2 precursor (Acrosin-trypsin inhibitor) (Epididymis tissue protein Li 172) (HUSI-II)
SPOCK3        4q32.3          Q9BQ16     TICN3_HUMAN 607989 Testican-3 precursor (SPARC/osteonectin, CWCV, and Kazal-like domains proteoglycan 3) [TICN3] [UNQ409/PRO771]
SPON2         4p16.3          Q9BUD6     SPON2_HUMAN 605918 Spondin-2 precursor (Differentially expressed in cancerous and non-cancerous lung cells 1) (DIL-1) (Mindin) [DIL1] [UNQ435/PRO866]
SPP1          4q21-q25        P10451     OSTP_HUMAN  166490 Osteopontin precursor (Bone sialoprotein 1) (Nephropontin) (Secreted phosphoprotein 1) (SPP-1) (Urinary stone protein) (Uropontin) [BNSP] [OPN] [PSEC0156]
SPRY1         4q26            O43609     SPY1_HUMAN  602465 Protein sprouty homolog 1 (Spry-1)
SRD5A3        4q12            Q9H8P0     PORED_HUMAN 611715 Polyprenol reductase (EC (3-oxo-5-alpha-steroid 4-dehydrogenase 3) (EC (Steroid 5-alpha-reductase 2-like) (Steroid 5-alpha-reductase 3) (S5AR 3) (SR type 3) [SRD5A2L]
SRP72         4q11            O76094     SRP72_HUMAN 602122 Signal recognition particle subunit SRP72 (SRP72) (Signal recognition particle 72 kDa protein)
STAP1         4q13.2          Q9ULZ2     STAP1_HUMAN 604298 Signal-transducing adaptor protein 1 (STAP-1) (BCR downstream-signaling protein 1) (Docking protein BRDG1) (Stem cell adaptor protein 1) [BRDG1]
STATH         4q11-q13        P02808     STAT_HUMAN  184470 Statherin precursor
STBD1         4q24-q25        O95210     STBD1_HUMAN 607406 Starch-binding domain-containing protein 1 (Genethonin-1) (Glycophagy cargo receptor STBD1) [GENX-3414]
STIM2         4p15.1          Q9P246     STIM2_HUMAN 610841 Stromal interaction molecule 2 precursor [KIAA1482]
STK32B        4p16            Q9NY57     ST32B_HUMAN        Serine/threonine-protein kinase 32B (EC (Yet another novel kinase 2) [YANK2] [UNQ3003/PRO9744]
STOX2         4q35.1          Q9P2F5     STOX2_HUMAN 617359 Storkhead-box protein 2 [KIAA1392]
STPG2         4q22.3-q23      Q8N412     STPG2_HUMAN        Sperm-tail PG-rich repeat-containing protein 2 [C4orf37]
STX18         4p16.2-p16.3    Q9P2W9     STX18_HUMAN 606046 Syntaxin-18 (Cell growth-inhibiting gene 9 protein) [GIG9]
SULT1B1       4q13.3          O43704     ST1B1_HUMAN 608436 Sulfotransferase 1B1 (EC (ST1B1) (Sulfotransferase 1B2) (Sulfotransferase family cytosolic 1B member 1) (Thyroid hormone sulfotransferase) [ST1B2] [SULT1B2]
SULT1E1       4q13.1          P49888     ST1E1_HUMAN 600043 Sulfotransferase 1E1 (EC (ST1E1) (EST-1) (Estrogen sulfotransferase) (Sulfotransferase, estrogen-preferring) [STE]
SYNPO2        4q26            Q9UMS6     SYNP2_HUMAN        Synaptopodin-2 (Genethonin-2) (Myopodin)
SYT14P1       4q13.2          Q58G82     SY14L_HUMAN 610892 Putative synaptotagmin-14-like protein (Synaptotagmin XIV protein pseudogene 1) (Synaptotagmin XIV-derived protein) (Synaptotagmin XIV-like protein) [SYT14L] [SYT14LP1] [SYTDEP]
TACC3         4p16.3          Q9Y6A5     TACC3_HUMAN 605303 Transforming acidic coiled-coil-containing protein 3 (ERIC-1) [ERIC1]
TACR3         4q25            P29371     NK3R_HUMAN  162332 Neuromedin-K receptor (NKR) (NK-3 receptor) (NK-3R) (Neurokinin B receptor) (Tachykinin receptor 3) [NK3R] [TAC3R]
TADA2B        4p16.1          Q86TJ2     TAD2B_HUMAN 608790 Transcriptional adapter 2-beta (ADA2-like protein beta) (ADA2-beta) [ADA2B]
TAPT1         4p15.32         Q6NXT6     TAPT1_HUMAN 612758 Transmembrane anterior posterior transformation protein 1 homolog (Cytomegalovirus partial fusion receptor) [CMVFR]
TBC1D1        4p14            Q86TI0     TBCD1_HUMAN 609850 TBC1 domain family member 1 [KIAA1108]
TBC1D14       4p16.1          Q9P2M4     TBC14_HUMAN 614855 TBC1 domain family member 14 [KIAA1322]
TBC1D19       4p15.2          Q8N5T2     TBC19_HUMAN        TBC1 domain family member 19
TBC1D9        4q31.21         Q6ZT07     TBCD9_HUMAN 618035 TBC1 domain family member 9 (TBC1 domain family member 9A) [KIAA0882] [TBC1D9A]
TBCK          4q24            Q8TEA7     TBCK_HUMAN  616899 TBC domain-containing protein kinase-like protein [TBCKL] [HSPC302]
TDO2          4q31-q32        P48775     T23O_HUMAN  191070 Tryptophan 2,3-dioxygenase (EC (TDO) (Tryptamin 2,3-dioxygenase) (Tryptophan oxygenase) (TO) (TRPO) (Tryptophan pyrrolase) (Tryptophanase) [TDO]
TEC           4p12            P42680     TEC_HUMAN   600583 Tyrosine-protein kinase Tec (EC [PSCTK4]
TECRL         4q13.1          Q5HYJ1     TECRL_HUMAN 617242 Trans-2,3-enoyl-CoA reductase-like (EC 1.3.1.-) (Steroid 5-alpha-reductase 2-like 2 protein) [SRD5A2L2]
TENM3         4q35.1          Q9P273     TEN3_HUMAN  610083 Teneurin-3 (Ten-3) (Protein Odd Oz/ten-m homolog 3) (Tenascin-M3) (Ten-m3) (Teneurin transmembrane protein 3) [KIAA1455] [ODZ3] [TNM3]
TET2          4q24            Q6N021     TET2_HUMAN  612839 Methylcytosine dioxygenase TET2 (EC 1.14.11.n2) [KIAA1546] [Nbla00191]
THAP6         4q21.21         Q8TBB0     THAP6_HUMAN 612535 THAP domain-containing protein 6
THAP9         4q21.22         Q9H5L6     THAP9_HUMAN 612537 DNA transposase THAP9 (EC 2.7.7.-) (THAP domain-containing protein 9) (hTh9)
THEGL         4q12            P0DJG4     THEGL_HUMAN        Testicular haploid expressed gene protein-like (Theg spermatid-like protein)
TIFA          4q25            Q96CG3     TIFA_HUMAN  609028 TRAF-interacting protein with FHA domain-containing protein A (Putative MAPK-activating protein PM14) (Putative NF-kappa-B-activating protein 20) (TRAF2-binding protein) [T2BP]
TIGD2         4q22.1          Q4W5G0     TIGD2_HUMAN 612973 Tigger transposable element-derived protein 2
TIGD4         4q31.3          Q8IY51     TIGD4_HUMAN        Tigger transposable element-derived protein 4
TKTL2         4q32.2          Q9H0I9     TKTL2_HUMAN        Transketolase-like protein 2 (EC
TLL1          4q32-q33        O43897     TLL1_HUMAN  606742 Tolloid-like protein 1 precursor (EC 3.4.24.-) [TLL]
TLR1          4p14            Q15399     TLR1_HUMAN  601194 Toll-like receptor 1 precursor (EC (Toll/interleukin-1 receptor-like protein) (TIL) (CD281 antigen) [KIAA0012]
TLR10         4p14            Q9BXR5     TLR10_HUMAN 606270 Toll-like receptor 10 precursor (EC (CD290 antigen) [UNQ315/PRO358]
TLR2          4q32            O60603     TLR2_HUMAN  603028 Toll-like receptor 2 precursor (EC (Toll/interleukin-1 receptor-like protein 4) (CD282 antigen) [TIL4]
TLR3          4q35            O15455     TLR3_HUMAN  603029 Toll-like receptor 3 precursor (CD283 antigen)
TLR6          4p16.1          Q9Y2C9     TLR6_HUMAN  605403 Toll-like receptor 6 precursor (EC (CD286 antigen)
TMA16         4q32.3          Q96EY4     TMA16_HUMAN        Translation machinery-associated protein 16 [C4orf43]
TMEM128       4p16.2          Q5BJH2     TM128_HUMAN        Transmembrane protein 128
TMEM129       4p16.3          A0AVI4     TM129_HUMAN 615975 E3 ubiquitin-protein ligase TM129 (EC (RING-type E3 ubiquitin transferase TM129)
TMEM131L      4q31.3          A2VDJ0     T131L_HUMAN 616243 Transmembrane protein 131-like precursor [KIAA0922]
TMEM144       4q32.1          Q7Z5S9     TM144_HUMAN        Transmembrane protein 144
TMEM150C      4q21.22         B9EJG8     T150C_HUMAN        Transmembrane protein 150C
TMEM154       4q31.3          Q6P9G4     TM154_HUMAN        Transmembrane protein 154 precursor
TMEM156       4p14            Q8N614     TM156_HUMAN        Transmembrane protein 156
TMEM165       4q12            Q9HC07     TM165_HUMAN 614726 Transmembrane protein 165 precursor (Transmembrane protein PT27) (Transmembrane protein TPARL) [TPARL]
TMEM175       4p16.3          Q9BSA9     TM175_HUMAN 616660 Endosomal/lysosomal potassium channel TMEM175 (Transmembrane protein 175) (hTMEM175)
TMEM184C      4q31.22         Q9NVA4     T184C_HUMAN 613937 Transmembrane protein 184C (Transmembrane protein 34) [TMEM34] [PRO1355]
TMEM192       4q32.3          Q8IY95     TM192_HUMAN        Transmembrane protein 192
TMEM271       4p16.3          A0A286YF58 TM271_HUMAN        Transmembrane protein 271
TMEM33        4p14            P57088     TMM33_HUMAN 618515 Transmembrane protein 33 (Protein DB83) (SHINC-3) [DB83]
TMPRSS11A     4q13.2          Q6ZMR5     TM11A_HUMAN 611704 Transmembrane protease serine 11A (EC 3.4.21.-) (Airway trypsin-like protease 1) (Epidermal type-II transmembrane serine protease) (Esophageal cancer-susceptibility gene 1 protein) [ECRG1] [HATL1] [HESP]
TMPRSS11B     4q13.2          Q86T26     TM11B_HUMAN        Transmembrane protease serine 11B (EC 3.4.21.-) (Airway trypsin-like protease 5) [HATL5]
TMPRSS11D     4q13.3          O60235     TM11D_HUMAN 605369 Transmembrane protease serine 11D precursor (EC 3.4.21.-) (Airway trypsin-like protease) [Contains: Transmembrane protease serine 11D non-catalytic chain; Transmembrane protease serine 11D catalytic chain] [HAT]
TMPRSS11E     4q13.1          Q9UL52     TM11E_HUMAN 610399 Transmembrane protease serine 11E precursor (EC 3.4.21.-) (Serine protease DESC1) (Transmembrane protease serine 11E2) [Contains: Transmembrane protease serine 11E non-catalytic chain; Transmembrane protease serine 11E catalytic chain] [DESC1] [TMPRSS11E2] [UNQ742/PRO1461]
TMPRSS11F     4q13.2          Q6ZWK6     TM11F_HUMAN        Transmembrane protease serine 11F (EC 3.4.21.-) (Airway trypsin-like protease 4) [HATL4]
TNIP2         4p16.3          Q8NFZ5     TNIP2_HUMAN 610669 TNFAIP3-interacting protein 2 (A20-binding inhibitor of NF-kappa-B activation 2) (ABIN-2) (Fetal liver LKB1-interacting protein) [ABIN2] [FLIP1]
TNIP3         4q27            Q96KP6     TNIP3_HUMAN 608019 TNFAIP3-interacting protein 3 (A20-binding inhibitor of NF-kappa-B activation 3) (ABIN-3) (Listeria-induced gene protein) [ABIN3] [LIND]
TRAM1L1       4q26            Q8N609     TR1L1_HUMAN 617505 Translocating chain-associated membrane protein 1-like 1
TRAPPC11      4q35.1          Q7Z392     TPC11_HUMAN 614138 Trafficking protein particle complex subunit 11 [C4orf41]
TRIM2         4q31.23         Q9C040     TRIM2_HUMAN 614141 Tripartite motif-containing protein 2 (EC (E3 ubiquitin-protein ligase TRIM2) (RING finger protein 86) (RING-type E3 ubiquitin transferase TRIM2) [KIAA0517] [RNF86]
TRIM60        4q32.3          Q495X7     TRI60_HUMAN        Tripartite motif-containing protein 60 (RING finger protein 129) (RING finger protein 33) [RNF129] [RNF33]
TRIM61        4q32.3          Q5EBN2     TRI61_HUMAN        Putative tripartite motif-containing protein 61 (RING finger protein 35) [RNF35]
TRIM75P       4q32.3          A6NK02     TRI75_HUMAN        Putative tripartite motif-containing protein 75 (Tripartite motif-containing protein 75 pseudogene) [TRIM75]
TRIML1        4q35.2          Q8N9V2     TRIML_HUMAN        Probable E3 ubiquitin-protein ligase TRIML1 (EC (RING finger protein 209) (RING-type E3 ubiquitin transferase TRIML1) (Tripartite motif family-like protein 1) [RNF209]
TRIML2        4q35.2          Q8N7C3     TRIMM_HUMAN        Probable E3 ubiquitin-protein ligase TRIML2 (EC (RING-type E3 ubiquitin transferase TRIML2) (SPRY domain-containing protein 6) (Tripartite motif family-like protein 2) [SPRYD6]
TRMT10A       4q23            Q8TBZ6     TM10A_HUMAN 616013 tRNA methyltransferase 10 homolog A (EC (RNA (guanine-9-)-methyltransferase domain-containing protein 2) (tRNA (guanine(9)-N(1))-methyltransferase TRMT10A) [RG9MTD2]
TRMT44        4p16.1          Q8IYL2     TRM44_HUMAN 614309 Probable tRNA (uracil-O(2)-)-methyltransferase (EC (Methyltransferase-like protein 19) [C4orf23] [METTL19]
TRPC3         4q28.1          Q13507     TRPC3_HUMAN 602345 Short transient receptor potential channel 3 (TrpC3) (Transient receptor protein 3) (TRP-3) (hTrp-3) (hTrp3) [TRP3]
TSPAN5        4p23            P62079     TSN5_HUMAN  613136 Tetraspanin-5 (Tspan-5) (Tetraspan NET-4) (Transmembrane 4 superfamily member 9) [TM4SF9]
TTC29         4q31.23         Q8NA56     TTC29_HUMAN 618735 Tetratricopeptide repeat protein 29 (TPR repeat protein 29) (Protein TBPP2A) (Testis development protein NYD-SP14)
TXK           4p12            P42681     TXK_HUMAN   600058 Tyrosine-protein kinase TXK (EC (Protein-tyrosine kinase 4) (Resting lymphocyte kinase) [PTK4] [RLK]
UBA6          4q13.2          A0AVT1     UBA6_HUMAN  611361 Ubiquitin-like modifier-activating enzyme 6 (EC (Ubiquitin-activating enzyme 6) (Monocyte protein 4) (MOP-4) (Ubiquitin-activating enzyme E1-like protein 2) (E1-L2) [MOP4] [UBE1L2]
UBE2D3        4q22.2          P61077     UB2D3_HUMAN 602963 Ubiquitin-conjugating enzyme E2 D3 (EC ((E3-independent) E2 ubiquitin-conjugating enzyme D3) (EC (E2 ubiquitin-conjugating enzyme D3) (Ubiquitin carrier protein D3) (Ubiquitin-conjugating enzyme E2(17)KB 3) (Ubiquitin-conjugating enzyme E2-17 kDa 3) (Ubiquitin-protein ligase D3) [UBC5C] [UBCH5C]
UBE2K         4p14            P61086     UBE2K_HUMAN 602846 Ubiquitin-conjugating enzyme E2 K (EC (E2 ubiquitin-conjugating enzyme K) (Huntingtin-interacting protein 2) (HIP-2) (Ubiquitin carrier protein) (Ubiquitin-conjugating enzyme E2-25 kDa) (Ubiquitin-conjugating enzyme E2(25K)) (Ubiquitin-conjugating enzyme E2-25K) (Ubiquitin-protein ligase) [HIP2] [LIG]
UCHL1         4p14            P09936     UCHL1_HUMAN 191342 Ubiquitin carboxyl-terminal hydrolase isozyme L1 precursor (EC (UCH-L1) (Neuron cytoplasmic protein 9.5) (PGP 9.5) (PGP9.5) (Ubiquitin thioesterase L1)
UCP1          4q28-q31        P25874     UCP1_HUMAN  113730 Mitochondrial brown fat uncoupling protein 1 (UCP 1) (Solute carrier family 25 member 7) (Thermogenin) [SLC25A7] [UCP]
UFSP2         4q35.1          Q9NUQ7     UFSP2_HUMAN 611482 Ufm1-specific protease 2 (EC 3.4.22.-) (UfSP2) [C4orf20]
UGDH          4p15.1          O60701     UGDH_HUMAN  603370 UDP-glucose 6-dehydrogenase (EC (UDP-Glc dehydrogenase) (UDP-GlcDH) (UDPGDH)
UGT2A1        4q13            P0DTE4     UD2A1_HUMAN 604716 UDP-glucuronosyltransferase 2A1 precursor (EC (UDPGT 2A1) (UGT2A1)
UGT2A2        4q13            P0DTE5     UD2A2_HUMAN 604716 UDP-glucuronosyltransferase 2A2 (EC (UDPGT 2A2)
UGT2A3        4q13.2          Q6UWM9     UD2A3_HUMAN 616382 UDP-glucuronosyltransferase 2A3 precursor (EC (UDPGT 2A3) [UNQ2559/PRO6239]
UGT2B10       4q13.1          P36537     UDB10_HUMAN 600070 UDP-glucuronosyltransferase 2B10 precursor (EC (UDPGT 2B10)
UGT2B11       4q13.1          O75310     UDB11_HUMAN 603064 UDP-glucuronosyltransferase 2B11 precursor (EC (UDPGT 2B11)
UGT2B15       4q13            P54855     UDB15_HUMAN 600069 UDP-glucuronosyltransferase 2B15 precursor (EC (UDPGT 2B15) (UGT2B15) (HLUG4) (UDP-glucuronosyltransferase 2B8) (UDPGT 2B8) (UDPGTh-3) [UGT2B8]
UGT2B17       4q13            O75795     UDB17_HUMAN 601903 UDP-glucuronosyltransferase 2B17 precursor (EC (UDPGT 2B17) (UGT2B17) (C19-steroid-specific UDP-glucuronosyltransferase) (C19-steroid-specific UDPGT)
UGT2B28       4q13.2          Q9BY64     UDB28_HUMAN 606497 UDP-glucuronosyltransferase 2B28 precursor (EC (UDPGT 2B28) (UGT2B28)
UGT2B4        4q13            P06133     UD2B4_HUMAN 600067 UDP-glucuronosyltransferase 2B4 precursor (EC (UDPGT 2B4) (UGT2B4) (HLUG25) (Hyodeoxycholic acid-specific UDPGT) (UDPGTh-1) [UGT2B11]
UGT2B7        4q13            P16662     UD2B7_HUMAN 600068 UDP-glucuronosyltransferase 2B7 precursor (EC (UDPGT 2B7) (UGT2B7) (3,4-catechol estrogen-specific UDPGT) (UDP-glucuronosyltransferase 2B9) (UDPGT 2B9) (UDPGTh-2) [UGTB2B9]
UGT8          4q26            Q16880     CGT_HUMAN   601291 2-hydroxyacylsphingosine 1-beta-galactosyltransferase precursor (EC (Ceramide UDP-galactosyltransferase) (Cerebroside synthase) (UDP-galactose-ceramide galactosyltransferase) [CGT] [UGT4]
UNC5C         4q21-q23        O95185     UNC5C_HUMAN 603610 Netrin receptor UNC5C precursor (Protein unc-5 homolog 3) (Protein unc-5 homolog C) [UNC5H3]
USO1          4q21.1-q21.3    O60763     USO1_HUMAN  603344 General vesicular transport factor p115 (Protein USO1 homolog) (Transcytosis-associated protein) (TAP) (Vesicle-docking protein) [VDP]
USP17L10      4p16.1          C9JJH3     U17LA_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 10 (EC
USP17L11      4p16.1          C9JVI0     U17LB_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 11 (EC
USP17L12      4p16.1          C9JPN9     UL17C_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 12 (EC
USP17L13      4p16.1          C9JLJ4     U17LD_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 13 (EC
USP17L15      4p16.1          C9J2P7     U17LF_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 15 (EC
USP17L17      4p16.1          D6RBQ6     U17LH_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 17 (EC
USP17L18      4p16.1          D6R9N7     U17LI_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 18 (EC
USP17L19      4p16.1          D6RCP7     U17LJ_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 19 (EC
USP17L20      4p16.1          D6RJB6     U17LK_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 20 (EC
USP17L21      4p16.1          D6R901     U17LL_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 21 (EC
USP17L22      4p16.1          D6RA61     U17LM_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 22 (EC
USP17L23      4p16.1          D6RBM5     U17LN_HUMAN        Putative ubiquitin carboxyl-terminal hydrolase 17-like protein 23
USP17L24      4p16.1          Q0WX57     U17LO_HUMAN 607011 Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17) [USP17] [USP17H] [USP17I] [USP17J] [USP17K] [USP17L] [USP17M]
USP17L25      4p16.1          Q0WX57     U17LO_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17)
USP17L26      4p16.1          Q0WX57     U17LO_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17)
USP17L27      4p16.1          Q0WX57     U17LO_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17)
USP17L28      4p16.1          Q0WX57     U17LO_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17)
USP17L29      4p16.1          Q0WX57     U17LO_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17)
USP17L30      4p16.1          Q0WX57     U17LO_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 24 (EC (Deubiquitinating enzyme 17) (Ubiquitin thioesterase 17) (Ubiquitin-specific-processing protease 17)
USP17L5       4p16.1          A8MUK1     U17L5_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 5 (EC (Deubiquitinating enzyme 17-like protein 5) (Ubiquitin thioesterase 17-like protein 5) (Ubiquitin-specific-processing protease 17-like protein 5)
USP17L6P      4p16.1          Q6QN14     U17L6_HUMAN        Ubiquitin carboxyl-terminal hydrolase 17-like protein 6 (EC (Deubiquitinating enzyme 17-like protein 6) (Ubiquitin thioesterase 17-like protein 6) (Ubiquitin-specific-processing protease 17-like protein 6) [USP17C] [USP17D] [USP17N]
USP38         4q31.1          Q8NB14     UBP38_HUMAN 618322 Ubiquitin carboxyl-terminal hydrolase 38 (EC (Deubiquitinating enzyme 38) (HP43.8KD) (Ubiquitin thioesterase 38) (Ubiquitin-specific-processing protease 38) [KIAA1891]
USP46         4q12            P62068     UBP46_HUMAN 612849 Ubiquitin carboxyl-terminal hydrolase 46 (EC (Deubiquitinating enzyme 46) (Ubiquitin thioesterase 46) (Ubiquitin-specific-processing protease 46)
USP53         4q26            Q70EK8     UBP53_HUMAN 617431 Inactive ubiquitin carboxyl-terminal hydrolase 53 (Inactive ubiquitin-specific peptidase 53) [KIAA1350]
UTP3          4q13.3          Q9NQZ2     SAS10_HUMAN 611614 Something about silencing protein 10 (Charged amino acid-rich leucine zipper 1) (CRL1) (Disrupter of silencing SAS10) (UTP3 homolog) [CRLZ1] [SAS10]
UVSSA         4p16.3          Q2YD98     UVSSA_HUMAN 614632 UV-stimulated scaffold protein A [KIAA1530]
VEGFC         4q34.1-q34.3    P49767     VEGFC_HUMAN 601528 Vascular endothelial growth factor C precursor (VEGF-C) (Flt4 ligand) (Flt4-L) (Vascular endothelial growth factor-related protein) (VRP)
WDFY3         4q21.23         Q8IZQ1     WDFY3_HUMAN 617485 WD repeat and FYVE domain-containing protein 3 (Autophagy-linked FYVE protein) (Alfy) [KIAA0993]
WDR1          4p15.32         O75083     WDR1_HUMAN  604734 WD repeat-containing protein 1 (Actin-interacting protein 1) (AIP1) (NORI-1)
WDR17         4q34            Q8IZU2     WDR17_HUMAN 609005 WD repeat-containing protein 17
WDR19         4p14            Q8NEZ3     WDR19_HUMAN 608151 WD repeat-containing protein 19 (Intraflagellar transport 144 homolog) [IFT144] [KIAA1638]
WFS1          4p16.1          O76024     WFS1_HUMAN  606201 Wolframin
WWC2          4q35.1          Q6AWC2     WWC2_HUMAN         Protein WWC2 (BH-3-only member B) (WW domain-containing protein 2) [BOMB]
WWC2-AS2      4q35.1          Q96NR7     WWAS2_HUMAN        Putative uncharacterized protein WWC2-AS2 (WWC2 antisense RNA 2) (WWC2 antisense gene protein 2) [C4orf38]
YIPF7         4p13            Q8N8F6     YIPF7_HUMAN        Protein YIPF7 (Five-pass transmembrane protein localizing in the Golgi apparatus and the endoplasmic reticulum 9) (YIP1 family member 7) [FINGER9] [YIP1B]
YTHDC1        4q13.3          Q96MU7     YTDC1_HUMAN        YTH domain-containing protein 1 (Splicing factor YT521) (YT521-B) [KIAA1966] [YT521]
ZAR1          4p12            Q86SH2     ZAR1_HUMAN  607520 Zygote arrest protein 1 (Oocyte-specific maternal effect factor)
ZBTB49        4p16.3          Q6ZSB9     ZBT49_HUMAN 616238 Zinc finger and BTB domain-containing protein 49 (Zinc finger protein 509) [ZNF509]
ZCCHC4        4p15.2          Q9H5U6     ZCHC4_HUMAN 611792 rRNA N6-adenosine-methyltransferase ZCCHC4 (EC 2.1.1.-) (Zinc finger CCHC domain-containing protein 4)
ZFP42         4q35.2          Q96MM3     ZFP42_HUMAN 614572 Zinc finger protein 42 homolog (Zfp-42) (Reduced expression protein 1) (REX-1) (hREX-1) (Zinc finger protein 754) [REX1] [ZNF754]
ZFYVE28       4p16.3          Q9HCC9     LST2_HUMAN  614176 Lateral signaling target protein 2 homolog (hLst2) (Zinc finger FYVE domain-containing protein 28) [KIAA1643] [LST2]
ZGRF1         4q25            Q86YA3     ZGRF1_HUMAN        Protein ZGRF1 (GRF-type zinc finger domain-containing protein 1) [C4orf21]
ZNF141        4p16.3          Q15928     ZN141_HUMAN 194648 Zinc finger protein 141 [D4S90]
ZNF330        4q31.1-q31.2    Q9Y3S2     ZN330_HUMAN 609550 Zinc finger protein 330 (Nucleolar autoantigen 36) (Nucleolar cysteine-rich protein) [NOA36]
ZNF518B       4p16.1          Q9C0D4     Z518B_HUMAN 617734 Zinc finger protein 518B [KIAA1729]
ZNF595        4p16.3          Q8IYB9     ZN595_HUMAN        Zinc finger protein 595
ZNF718        4p16.3          Q3SXZ3     ZN718_HUMAN 618405 Zinc finger protein 718
ZNF721        4p16.3          Q8TF20     ZN721_HUMAN        Zinc finger protein 721 [KIAA1982]
ZNF732        4p16.3          B4DXR9     ZN732_HUMAN        Zinc finger protein 732
ZNF827        4q31.22         Q17R98     ZN827_HUMAN 617962 Zinc finger protein 827
ZNF876P       4p16.3          Q49A33     Z876P_HUMAN        Putative zinc finger protein 876
-             4p13-q14        Q8IZM0     CNG1O_HUMAN        Putative CNGA1-overlapping antisense gene protein (Anti-CNG1)
-             4p14            Q9H8V8     YD018_HUMAN        Putative uncharacterized protein FLJ13197
-             4p15.2          Q6ZS92     YD022_HUMAN        Putative uncharacterized protein FLJ45721
-             4p15.32         A8MVM7     YD021_HUMAN        Putative uncharacterized protein ENSP00000382790
-             4p16.1          B2RBV5     MRUPP_HUMAN        Putative MORF4 family-associated protein 1-like protein UPP (Unnamed protein product) (UPP)
-             4q35.1          Q6ZQT0     YD023_HUMAN        Putative uncharacterized protein FLJ45035
-             4q35.1          Q6ZRP5     YD019_HUMAN        Putative uncharacterized protein FLJ46204

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