##gff-version 3 Q5VUA4 UniProtKB Chain 1 2279 . . . ID=PRO_0000191807;Note=Zinc finger protein 318 Q5VUA4 UniProtKB Zinc finger 1063 1097 . . . Note=Matrin-type 1 Q5VUA4 UniProtKB Zinc finger 1136 1166 . . . Note=Matrin-type 2 Q5VUA4 UniProtKB Region 1 1092 . . . Note=Interaction with AR;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q99PP2 Q5VUA4 UniProtKB Region 1 140 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 164 189 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 279 346 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 397 416 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 514 533 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 540 570 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 664 709 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 922 942 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 989 1051 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 1195 1319 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 1428 1463 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 1577 1628 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 1702 1735 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 1753 1776 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Region 2252 2279 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Coiled coil 315 343 . . . Ontology_term=ECO:0000255;evidence=ECO:0000255 Q5VUA4 UniProtKB Coiled coil 876 980 . . . Ontology_term=ECO:0000255;evidence=ECO:0000255 Q5VUA4 UniProtKB Coiled coil 1768 1792 . . . Ontology_term=ECO:0000255;evidence=ECO:0000255 Q5VUA4 UniProtKB Compositional bias 75 99 . . . Note=Pro residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 113 132 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 164 180 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 286 301 . . . Note=Polar residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 305 336 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 397 413 . . . Note=Polar residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 519 533 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 664 689 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 990 1011 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 1012 1026 . . . Note=Polar residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 1195 1268 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 1277 1292 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 1299 1319 . . . Note=Basic and acidic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 1437 1462 . . . Note=Pro residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 1595 1627 . . . Note=Polar residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Compositional bias 2258 2279 . . . Note=Polar residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q5VUA4 UniProtKB Modified residue 40 40 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 79 79 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:21406692;Dbxref=PMID:21406692 Q5VUA4 UniProtKB Modified residue 81 81 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:21406692;Dbxref=PMID:21406692 Q5VUA4 UniProtKB Modified residue 136 136 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:21406692,ECO:0007744|PubMed:23186163;Dbxref=PMID:18669648,PMID:21406692,PMID:23186163 Q5VUA4 UniProtKB Modified residue 173 173 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:19690332,ECO:0007744|PubMed:24275569;Dbxref=PMID:18669648,PMID:19690332,PMID:24275569 Q5VUA4 UniProtKB Modified residue 205 205 . . . Note=Phosphotyrosine;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q99PP2 Q5VUA4 UniProtKB Modified residue 207 207 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 214 214 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:19690332;Dbxref=PMID:18669648,PMID:19690332 Q5VUA4 UniProtKB Modified residue 464 464 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:20068231;Dbxref=PMID:20068231 Q5VUA4 UniProtKB Modified residue 472 472 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 501 501 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 527 527 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:20068231,ECO:0007744|PubMed:21406692,ECO:0007744|PubMed:23186163;Dbxref=PMID:20068231,PMID:21406692,PMID:23186163 Q5VUA4 UniProtKB Modified residue 842 842 . . . Note=Phosphothreonine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 1010 1010 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 1037 1037 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:21406692,ECO:0007744|PubMed:24275569;Dbxref=PMID:21406692,PMID:24275569 Q5VUA4 UniProtKB Modified residue 1243 1243 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:17081983,ECO:0007744|PubMed:20068231,ECO:0007744|PubMed:21406692,ECO:0007744|PubMed:23186163;Dbxref=PMID:17081983,PMID:20068231,PMID:21406692,PMID:23186163 Q5VUA4 UniProtKB Modified residue 1267 1267 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:23186163,ECO:0007744|PubMed:24275569;Dbxref=PMID:23186163,PMID:24275569 Q5VUA4 UniProtKB Modified residue 1420 1420 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:20068231,ECO:0007744|PubMed:23186163;Dbxref=PMID:20068231,PMID:23186163 Q5VUA4 UniProtKB Modified residue 1713 1713 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:19690332;Dbxref=PMID:19690332 Q5VUA4 UniProtKB Modified residue 1856 1856 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 1896 1896 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:20068231,ECO:0007744|PubMed:23186163;Dbxref=PMID:18669648,PMID:20068231,PMID:23186163 Q5VUA4 UniProtKB Modified residue 1971 1971 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 2030 2030 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 2035 2035 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 2091 2091 . . . Note=Phosphoserine;Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:23186163;Dbxref=PMID:23186163 Q5VUA4 UniProtKB Modified residue 2101 2101 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:20068231,ECO:0007744|PubMed:21406692,ECO:0007744|PubMed:23186163,ECO:0007744|PubMed:24275569;Dbxref=PMID:20068231,PMID:21406692,PMID:23186163,PMID:24275569 Q5VUA4 UniProtKB Modified residue 2189 2189 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:23186163;Dbxref=PMID:18669648,PMID:23186163 Q5VUA4 UniProtKB Modified residue 2192 2192 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:23186163;Dbxref=PMID:18669648,PMID:23186163 Q5VUA4 UniProtKB Modified residue 2243 2243 . . . Note=Phosphoserine;Ontology_term=ECO:0007744,ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:18669648,ECO:0007744|PubMed:20068231,ECO:0007744|PubMed:23186163;Dbxref=PMID:18669648,PMID:20068231,PMID:23186163 Q5VUA4 UniProtKB Cross-link 547 547 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2);Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:28112733;Dbxref=PMID:28112733 Q5VUA4 UniProtKB Cross-link 553 553 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2);Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:28112733;Dbxref=PMID:28112733 Q5VUA4 UniProtKB Cross-link 566 566 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2);Ontology_term=ECO:0007744;evidence=ECO:0007744|PubMed:28112733;Dbxref=PMID:28112733 Q5VUA4 UniProtKB Cross-link 578 578 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2);Ontology_term=ECO:0007744,ECO:0007744;evidence=ECO:0007744|PubMed:25772364,ECO:0007744|PubMed:28112733;Dbxref=PMID:25772364,PMID:28112733 Q5VUA4 UniProtKB Alternative sequence 1093 1117 . . . ID=VSP_016592;Note=In isoform 2. TLDPYNRPWASKTQSEAKQDAIKRT->GQFQKSSDFQKEGLQQTFLPPERQG;Ontology_term=ECO:0000303;evidence=ECO:0000303|PubMed:15489334;Dbxref=PMID:15489334 Q5VUA4 UniProtKB Alternative sequence 1118 2279 . . . ID=VSP_016593;Note=In isoform 2. Missing;Ontology_term=ECO:0000303;evidence=ECO:0000303|PubMed:15489334;Dbxref=PMID:15489334 Q5VUA4 UniProtKB Natural variant 407 407 . . . ID=VAR_053759;Note=S->I;Dbxref=dbSNP:rs34541323 Q5VUA4 UniProtKB Natural variant 812 812 . . . ID=VAR_036056;Note=In a breast cancer sample%3B somatic mutation. N->S;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16959974;Dbxref=dbSNP:rs141660717,PMID:16959974 Q5VUA4 UniProtKB Natural variant 870 870 . . . ID=VAR_053760;Note=L->V;Dbxref=dbSNP:rs9357410 Q5VUA4 UniProtKB Natural variant 1274 1274 . . . ID=VAR_036057;Note=In a breast cancer sample%3B somatic mutation. G->R;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16959974;Dbxref=PMID:16959974 Q5VUA4 UniProtKB Natural variant 1292 1292 . . . ID=VAR_053761;Note=T->I;Dbxref=dbSNP:rs10948072 Q5VUA4 UniProtKB Natural variant 1580 1580 . . . ID=VAR_053762;Note=A->T;Dbxref=dbSNP:rs3734684 Q5VUA4 UniProtKB Natural variant 1583 1583 . . . ID=VAR_053763;Note=T->I;Dbxref=dbSNP:rs36107018 Q5VUA4 UniProtKB Natural variant 1797 1797 . . . ID=VAR_053764;Note=V->A;Ontology_term=ECO:0000269;evidence=ECO:0000269|Ref.1;Dbxref=dbSNP:rs1459675 Q5VUA4 UniProtKB Sequence conflict 62 62 . . . Note=G->R;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q5VUA4 UniProtKB Sequence conflict 75 75 . . . Note=G->V;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q5VUA4 UniProtKB Sequence conflict 1034 1035 . . . Note=RT->CI;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q5VUA4 UniProtKB Sequence conflict 1259 1259 . . . Note=L->F;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q5VUA4 UniProtKB Sequence conflict 1349 1349 . . . Note=R->G;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q5VUA4 UniProtKB Sequence conflict 2160 2160 . . . Note=S->F;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q5VUA4 UniProtKB Sequence conflict 2173 2173 . . . Note=V->D;Ontology_term=ECO:0000305;evidence=ECO:0000305