##gff-version 3 Q16778 UniProtKB Initiator methionine 1 1 . . . Note=Removed;Ontology_term=ECO:0000250,ECO:0000269,ECO:0000269,ECO:0000269,ECO:0000269;evidence=ECO:0000250|UniProtKB:P23527,ECO:0000269|PubMed:11859126,ECO:0000269|PubMed:12860195,ECO:0000269|PubMed:15019208,ECO:0000269|PubMed:8620898;Dbxref=PMID:11859126,PMID:12860195,PMID:15019208,PMID:8620898 Q16778 UniProtKB Chain 2 126 . . . ID=PRO_0000071835;Note=Histone H2B type 2-E Q16778 UniProtKB Region 1 35 . . . Note=Disordered;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q16778 UniProtKB Compositional bias 17 35 . . . Note=Basic residues;Ontology_term=ECO:0000256;evidence=ECO:0000256|SAM:MobiDB-lite Q16778 UniProtKB Modified residue 2 2 . . . Note=N-acetylproline;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:P23527 Q16778 UniProtKB Modified residue 3 3 . . . Note=ADP-ribosyl glutamic acid;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27530147;Dbxref=PMID:27530147 Q16778 UniProtKB Modified residue 6 6 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 6 6 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 6 6 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000269,ECO:0000269;evidence=ECO:0000269|PubMed:16283522,ECO:0000269|PubMed:16627869;Dbxref=PMID:16283522,PMID:16627869 Q16778 UniProtKB Modified residue 6 6 . . . Note=N6-butyryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105113;Dbxref=PMID:27105113 Q16778 UniProtKB Modified residue 6 6 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 6 6 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 7 7 . . . Note=ADP-ribosylserine;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:34874266;Dbxref=PMID:34874266 Q16778 UniProtKB Modified residue 12 12 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 12 12 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16627869;Dbxref=PMID:16627869 Q16778 UniProtKB Modified residue 12 12 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 12 12 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 13 13 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 13 13 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000269,ECO:0000269;evidence=ECO:0000269|PubMed:16283522,ECO:0000269|PubMed:16627869;Dbxref=PMID:16283522,PMID:16627869 Q16778 UniProtKB Modified residue 13 13 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 15 15 . . . Note=Phosphoserine%3B by STK4/MST1;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:12757711;Dbxref=PMID:12757711 Q16778 UniProtKB Modified residue 16 16 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000269,ECO:0000269;evidence=ECO:0000269|PubMed:16283522,ECO:0000269|PubMed:16627869;Dbxref=PMID:16283522,PMID:16627869 Q16778 UniProtKB Modified residue 16 16 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 16 16 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 17 17 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 17 17 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16627869;Dbxref=PMID:16627869 Q16778 UniProtKB Modified residue 17 17 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 17 17 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 17 17 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 21 21 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 21 21 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 21 21 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000269,ECO:0000269;evidence=ECO:0000269|PubMed:16283522,ECO:0000269|PubMed:16627869;Dbxref=PMID:16283522,PMID:16627869 Q16778 UniProtKB Modified residue 21 21 . . . Note=N6-butyryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105113;Dbxref=PMID:27105113 Q16778 UniProtKB Modified residue 21 21 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 21 21 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 24 24 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 24 24 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:P33778 Q16778 UniProtKB Modified residue 24 24 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 24 24 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 25 25 . . . Note=N6-(2-hydroxyisobutyryl)lysine;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 35 35 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 35 35 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 35 35 . . . Note=N6-crotonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21925322;Dbxref=PMID:21925322 Q16778 UniProtKB Modified residue 35 35 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 35 35 . . . Note=N6-succinyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:22389435;Dbxref=PMID:22389435 Q16778 UniProtKB Modified residue 36 36 . . . Note=PolyADP-ribosyl glutamic acid;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q64475 Q16778 UniProtKB Modified residue 37 37 . . . Note=Phosphoserine%3B by AMPK;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q64524 Q16778 UniProtKB Modified residue 44 44 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 44 44 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 44 44 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 47 47 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 47 47 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 47 47 . . . Note=N6-methyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16627869;Dbxref=PMID:16627869 Q16778 UniProtKB Modified residue 58 58 . . . Note=N6%2CN6-dimethyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16627869;Dbxref=PMID:16627869 Q16778 UniProtKB Modified residue 58 58 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 80 80 . . . Note=Dimethylated arginine;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q96A08 Q16778 UniProtKB Modified residue 86 86 . . . Note=N6%2CN6%2CN6-trimethyllysine%3B alternate;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q96A08 Q16778 UniProtKB Modified residue 86 86 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 86 86 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 86 86 . . . Note=N6-acetyllysine%3B alternate;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q96A08 Q16778 UniProtKB Modified residue 86 86 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 87 87 . . . Note=Omega-N-methylarginine;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q96A08 Q16778 UniProtKB Modified residue 93 93 . . . Note=Omega-N-methylarginine;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q96A08 Q16778 UniProtKB Modified residue 109 109 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 109 109 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 109 109 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 109 109 . . . Note=N6-methyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:16627869;Dbxref=PMID:16627869 Q16778 UniProtKB Modified residue 116 116 . . . Note=Phosphothreonine;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q00729 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-malonyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:22389435;Dbxref=PMID:22389435 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-methylated lysine%3B alternate;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q00729 Q16778 UniProtKB Modified residue 117 117 . . . Note=N6-succinyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:22389435;Dbxref=PMID:22389435 Q16778 UniProtKB Modified residue 121 121 . . . Note=N6-(2-hydroxyisobutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:24681537;Dbxref=PMID:24681537 Q16778 UniProtKB Modified residue 121 121 . . . Note=N6-(beta-hydroxybutyryl)lysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:27105115;Dbxref=PMID:27105115 Q16778 UniProtKB Modified residue 121 121 . . . Note=N6-glutaryllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31542297;Dbxref=PMID:31542297 Q16778 UniProtKB Modified residue 121 121 . . . Note=N6-lactoyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:31645732;Dbxref=PMID:31645732 Q16778 UniProtKB Modified residue 121 121 . . . Note=N6-succinyllysine%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:22389435;Dbxref=PMID:22389435 Q16778 UniProtKB Glycosylation 113 113 . . . Note=O-linked (GlcNAc) serine;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:P62807 Q16778 UniProtKB Cross-link 6 6 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2)%3B alternate;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:P58876 Q16778 UniProtKB Cross-link 21 21 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in SUMO2)%3B alternate;Ontology_term=ECO:0000250;evidence=ECO:0000250|UniProtKB:Q5QNW6 Q16778 UniProtKB Cross-link 35 35 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in ubiquitin)%3B alternate;Ontology_term=ECO:0000269;evidence=ECO:0000269|PubMed:21726816;Dbxref=PMID:21726816 Q16778 UniProtKB Cross-link 121 121 . . . Note=Glycyl lysine isopeptide (Lys-Gly) (interchain with G-Cter in ubiquitin)%3B alternate;Ontology_term=ECO:0000269,ECO:0000269,ECO:0000269;evidence=ECO:0000269|PubMed:16307923,ECO:0000269|PubMed:16627869,ECO:0000269|PubMed:16713563;Dbxref=PMID:16307923,PMID:16627869,PMID:16713563 Q16778 UniProtKB Sequence conflict 5 5 . . . Note=A->S;Ontology_term=ECO:0000305;evidence=ECO:0000305 Q16778 UniProtKB Helix 39 49 . . . Ontology_term=ECO:0007829;evidence=ECO:0007829|PDB:6KBB Q16778 UniProtKB Beta strand 54 56 . . . Ontology_term=ECO:0007829;evidence=ECO:0007829|PDB:6KBB Q16778 UniProtKB Helix 57 84 . . . Ontology_term=ECO:0007829;evidence=ECO:0007829|PDB:6KBB Q16778 UniProtKB Beta strand 88 90 . . . Ontology_term=ECO:0007829;evidence=ECO:0007829|PDB:6KBB Q16778 UniProtKB Helix 92 102 . . . Ontology_term=ECO:0007829;evidence=ECO:0007829|PDB:6KBB Q16778 UniProtKB Helix 105 123 . . . Ontology_term=ECO:0007829;evidence=ECO:0007829|PDB:6KBB