ID P2SAF_ARATH Reviewed; 403 AA. AC O82660; Q56Z72; DT 15-JUL-1999, integrated into UniProtKB/Swiss-Prot. DT 01-NOV-1998, sequence version 1. DT 16-JUN-2009, entry version 75. DE RecName: Full=Photosystem II stability/assembly factor HCF136, chloroplastic; DE Flags: Precursor; GN Name=HCF136; OrderedLocusNames=At5g23120; ORFNames=MYJ24.11; OS Arabidopsis thaliana (Mouse-ear cress). OC Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta; OC Spermatophyta; Magnoliophyta; eudicotyledons; core eudicotyledons; OC rosids; eurosids II; Brassicales; Brassicaceae; Arabidopsis. OX NCBI_TaxID=3702; RN [1] RP NUCLEOTIDE SEQUENCE [MRNA], FUNCTION, AND SUBCELLULAR LOCATION. RC STRAIN=cv. Wassilewskija; TISSUE=Leaf; RX MEDLINE=98409534; PubMed=9736608; DOI=10.1093/emboj/17.18.5286; RA Meurer J., Pluecken H., Kowallik K.V., Westhoff P.; RT "A nuclear-encoded protein of prokaryotic origin is essential for the RT stability of photosystem II in Arabidopsis thaliana."; RL EMBO J. 17:5286-5297(1998). RN [2] RP NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA]. RC STRAIN=cv. Columbia; RX MEDLINE=98069011; PubMed=9405937; DOI=10.1093/dnares/4.4.291; RA Kotani H., Nakamura Y., Sato S., Kaneko T., Asamizu E., Miyajima N., RA Tabata S.; RT "Structural analysis of Arabidopsis thaliana chromosome 5. II. RT Sequence features of the regions of 1,044,062 bp covered by thirteen RT physically assigned P1 clones."; RL DNA Res. 4:291-300(1997). RN [3] RP NUCLEOTIDE SEQUENCE [LARGE SCALE MRNA]. RC STRAIN=cv. Columbia; RX MEDLINE=22954850; PubMed=14593172; DOI=10.1126/science.1088305; RA Yamada K., Lim J., Dale J.M., Chen H., Shinn P., Palm C.J., RA Southwick A.M., Wu H.C., Kim C.J., Nguyen M., Pham P.K., Cheuk R.F., RA Karlin-Newmann G., Liu S.X., Lam B., Sakano H., Wu T., Yu G., RA Miranda M., Quach H.L., Tripp M., Chang C.H., Lee J.M., Toriumi M.J., RA Chan M.M., Tang C.C., Onodera C.S., Deng J.M., Akiyama K., Ansari Y., RA Arakawa T., Banh J., Banno F., Bowser L., Brooks S.Y., Carninci P., RA Chao Q., Choy N., Enju A., Goldsmith A.D., Gurjal M., Hansen N.F., RA Hayashizaki Y., Johnson-Hopson C., Hsuan V.W., Iida K., Karnes M., RA Khan S., Koesema E., Ishida J., Jiang P.X., Jones T., Kawai J., RA Kamiya A., Meyers C., Nakajima M., Narusaka M., Seki M., Sakurai T., RA Satou M., Tamse R., Vaysberg M., Wallender E.K., Wong C., Yamamura Y., RA Yuan S., Shinozaki K., Davis R.W., Theologis A., Ecker J.R.; RT "Empirical analysis of transcriptional activity in the Arabidopsis RT genome."; RL Science 302:842-846(2003). RN [4] RP NUCLEOTIDE SEQUENCE [LARGE SCALE MRNA] OF 333-403. RC STRAIN=cv. Columbia; RA Totoki Y., Seki M., Ishida J., Nakajima M., Enju A., Kamiya A., RA Narusaka M., Shin-i T., Nakagawa M., Sakamoto N., Oishi K., Kohara Y., RA Kobayashi M., Toyoda A., Sakaki Y., Sakurai T., Iida K., Akiyama K., RA Satou M., Toyoda T., Konagaya A., Carninci P., Kawai J., RA Hayashizaki Y., Shinozaki K.; RT "Large-scale analysis of RIKEN Arabidopsis full-length (RAFL) cDNAs."; RL Submitted (MAR-2005) to the EMBL/GenBank/DDBJ databases. RN [5] RP PROTEIN SEQUENCE OF N-TERMINUS, AND IDENTIFICATION BY MASS RP SPECTROMETRY. RX MEDLINE=21684138; PubMed=11826309; DOI=10.1105/tpc.010304; RA Peltier J.-B., Emanuelsson O., Kalume D.E., Ytterberg J., Friso G., RA Rudella A., Liberles D.A., Soederberg L., Roepstorff P., RA von Heijne G., van Wijk K.J.; RT "Central functions of the lumenal and peripheral thylakoid proteome of RT Arabidopsis determined by experimentation and genome-wide RT prediction."; RL Plant Cell 14:211-236(2002). RN [6] RP IN VITRO IMPORT INTO PEA CHLOROPLASTS, AND MUTAGENESIS OF RP 55-ARG-ARG-56. RX PubMed=10664464; DOI=10.1016/S0014-5793(00)01129-7; RA Hynds P.J., Pluecken H., Westhoff P., Robinson C.; RT "Different lumen-targeting pathways for nuclear-encoded versus RT cyanobacterial/plastid-encoded Hcf136 proteins."; RL FEBS Lett. 467:97-100(2000). RN [7] RP FUNCTION IN EARLY ASSEMBLY OF PHOTOSYSTEM II. RX PubMed=12459468; DOI=10.1016/S0014-5793(02)03634-7; RA Pluecken H., Mueller B., Grohmann D., Westhoff P., Eichacker L.A.; RT "The HCF136 protein is essential for assembly of the photosystem II RT reaction center in Arabidopsis thaliana."; RL FEBS Lett. 532:85-90(2002). RN [8] RP IDENTIFICATION BY MASS SPECTROMETRY. RX PubMed=18431481; DOI=10.1371/journal.pone.0001994; RA Zybailov B., Rutschow H., Friso G., Rudella A., Emanuelsson O., RA Sun Q., van Wijk K.J.; RT "Sorting signals, N-terminal modifications and abundance of the RT chloroplast proteome."; RL PLoS ONE 3:E1994-E1994(2008). CC -!- FUNCTION: Essential for photosystem II (PSII) biogenesis; required CC for assembly of an early intermediate in PSII assembly that CC includes D2 (psbD) and cytochrome b559. Has been suggested CC (PubMed:11826309) to be required for chlorophyll a binding. CC -!- SUBCELLULAR LOCATION: Plastid, chloroplast thylakoid membrane; CC Peripheral membrane protein; Lumenal side. Note=Restricted to the CC stromal lamelae. Translocation into the thylakoid lumen occurs via CC the Tat pathway. CC -!- TISSUE SPECIFICITY: Expression in green tissue, not roots. CC -!- DEVELOPMENTAL STAGE: Accumulates also in dark-grown seedlings. CC -!- SIMILARITY: Belongs to the ycf48 family. CC -!- SIMILARITY: Contains 4 BNR repeats. CC ----------------------------------------------------------------------- CC Copyrighted by the UniProt Consortium, see http://www.uniprot.org/terms CC Distributed under the Creative Commons Attribution-NoDerivs License CC ----------------------------------------------------------------------- DR EMBL; Y15628; CAA75723.1; -; mRNA. DR EMBL; AB006708; BAB09829.1; -; Genomic_DNA. DR EMBL; AY045691; AAK74049.1; -; mRNA. DR EMBL; BT004549; AAO42795.1; -; mRNA. DR EMBL; AK221097; BAD94976.1; -; mRNA. DR IPI; IPI00539527; -. DR PIR; T51828; T51828. DR RefSeq; NP_197703.1; -. DR UniGene; At.5360; -. DR PRIDE; O82660; -. DR ProMEX; O82660; -. DR GeneID; 832376; -. DR GenomeReviews; BA000015_GR; AT5G23120. DR KEGG; ath:AT5G23120; -. DR NMPDR; fig|3702.1.peg.24483; -. DR TAIR; At5g23120; -. DR OMA; O82660; GFLVGSN. DR ArrayExpress; O82660; -. DR GermOnline; AT5G23120; Arabidopsis thaliana. DR GO; GO:0009533; C:chloroplast stromal thylakoid; IDA:TAIR. DR GO; GO:0009535; C:chloroplast thylakoid membrane; IDA:TAIR. DR GO; GO:0009523; C:photosystem II; IEA:UniProtKB-KW. DR GO; GO:0031977; C:thylakoid lumen; IDA:TAIR. DR GO; GO:0005515; F:protein binding; IDA:TAIR. DR GO; GO:0015979; P:photosynthesis; IEA:UniProtKB-KW. DR GO; GO:0009657; P:plastid organization; IMP:TAIR. DR GO; GO:0006461; P:protein complex assembly; IMP:TAIR. DR GO; GO:0046686; P:response to cadmium ion; IEP:TAIR. DR InterPro; IPR016705; PSII_stabil/assmbl_fac_HCF136. DR InterPro; IPR006311; Tat. DR PIRSF; PIRSF017875; PSII_HCF136; 1. DR TIGRFAMs; TIGR01409; TAT_signal_seq; 1. PE 1: Evidence at protein level; KW Chloroplast; Complete proteome; Direct protein sequencing; Membrane; KW Photosynthesis; Photosystem II; Plastid; Repeat; Thylakoid; KW Transit peptide. FT TRANSIT 1 53 Chloroplast (Potential). FT TRANSIT 54 78 Thylakoid. FT CHAIN 79 403 Photosystem II stability/assembly factor FT HCF136, chloroplastic. FT /FTId=PRO_0000005341. FT REPEAT 121 132 BNR 1. FT REPEAT 167 178 BNR 2. FT REPEAT 210 221 BNR 3. FT REPEAT 356 367 BNR 4. FT MUTAGEN 55 56 RR->KK: Protein is imported into FT chloroplasts but is unable to translocate FT into the thylakoid lumen. SQ SEQUENCE 403 AA; 44104 MW; 11079552F817FF9D CRC64; MASLQLCDGY LLFKPSVSPR FLSQRISHRL IPKASSSPPP SPSPSSSSSS LSFSRRELLY QSAAVSLSLS SIVGPARADE QLSEWERVFL PIDPGVVLLD IAFVPDEPSR GFLLGTRQTL LETKDGGSTW NPRSIPSAEE EDFNYRFNSI SFKGKEGWII GKPAILLYTA DAGENWDRIP LSSQLPGDMV FIKATEDKSA EMVTDEGAIY VTSNRGYNWK AAIQETVSAT LNRTVSSGIS GASYYTGTFS AVNRSPDGRY VAVSSRGNFF LTWEPGQPYW QPHNRAVARR IQNMGWRADG GLWLLVRGGG LYLSKGTGIT EEFEEVPVQS RGFGILDVGY RSEEEAWAAG GSGILLRTRN GGKSWNRDKA ADNIAANLYA VKFVDDKKGF VLGNDGVLLR YVG //